The division first formed on 1 September 1941 in the
Oryol Military District at Voronezh, right alongside the
327th Rifle Division. Col. Nikolai Matveevich Makovchuk took command the same day and remained in that post until 7 February 1942. The division's basic order of battle was as follows: • 1110th Rifle Regiment • 1112th Rifle Regiment • 1114th Rifle Regiment • 895th Artillery Regiment In late October the division was assigned to
26th Reserve Army, which was forming up in the
Reserve of the Supreme High Command. At the beginning of December, the 329th was in the Moscow Defense Zone in the
60th Reserve Army before being reassigned to
5th Army in Western Front.
Battle of Moscow 5th Army went over to the counteroffensive on December 6. By the close of December 11 the 329th, along with the
108th,
144th, and
19th Rifle Divisions had pierced the defensive front of the German
252nd,
87th,
78th and (coincidentally)
329th Infantry Divisions, and had reached the area of Lokotnya and Kolyubakovo. Western Front issued further orders for a continuation of the offensive on December 13. As well,
Maj. Gen. Lev Dovator's 2nd Guards Cavalry Corps was moved to the 5th Army sector in the woods north of
Kubinka. After a series of quick blows the rifle divisions mentioned above, plus the
50th, went over to the attack. Due to strong enemy resistance and extensive fortifications the assault made little immediate headway. However, enough of a gap was forced in the German line on the boundary between the 329th and the 19th divisions that the cavalry was able to pass through along a forlorn and roadless forest sector, beginning a deep raid which would disrupt and demoralize the enemy; already on the next day 5th Army resumed its advance. On December 20, the 19th and 329th Rifle Divisions forced the
Ruza River and liberated three villages. But this river line had been fortified by the Germans, and 5th Army's forces had lost many men and much equipment over the course of its fighting pursuit, so it was necessary to call a halt. In January 1942, the division formed part of the assault group of
33rd Army, which was driving through the lines of Army Group Center towards
Vyasma. This objective was not quite reached, and the army was cut off behind German lines by counterattacks in early February. On the 8th, the division came under the command of
Col. Kornei Mikhailovich Andrusenko. By the beginning of May, Western Front was developing a plan to break through to these isolated pockets with major forces and to hold them as staging areas for further operations. On May 10, Colonel Andrusenko handed command of the division to Col. Nikolai Lavrentevich Soldatov, who would hold the post until the division was disbanded. On May 20, by Front order, the "Special Group of General Belov" was created and consisted of
Lt. Gen. Pavel Belov's 1st Guards Cavalry Corps, the 329th Rifle Division, units of the
4th Airborne Corps, and several large partisan detachments. However, the plan was never implemented, due to renewed German offensive action and the poor organization of the airlift operations that were intended to support Group Belov. Late winter snow and early spring mud had delayed the inevitable, but by the end of the month the Germans had reduced the remaining men of the 329th to small, barely organized groups. The division was carried under the Western Front reserves until June, but by then it was clear that not enough of these men would slip out of the forests and back to Soviet lines, and on August 22 the unit was officially disbanded. Colonel Andrusenko was one of those fortunate enough to escape, but his story nearly ended tragically as he was put on trial before the military tribunal of Western Front "for criminal inaction" during the encirclement battle. On April 6 he was deprived of his military rank and sentenced to death. He applied to the Military Collegium of the Supreme Court of the USSR for a pardon, and on May 4 his sentence was reduced to 10 years in the
Gulag without full loss of rank, although he was demoted to Major. The sentence was also deferred until after the end of the war, so he continued to serve at the front. While Andrusenko had a further run-in with the military authorities later that year, he went on to regain his rank, be awarded the Gold Star
Hero of the Soviet Union for his leadership in crossing the
Dniepr, and take command of the
55th Rifle Division. ==2nd Formation==