He was active in many branches of botany, including anatomy and the taxonomy of
seed-producing plants. Among his achievements in this direction, the most notable is the treatise '''' ("On the generation and development of the spermatophyte embryo"), which is remarkable for the first account of any value of the development and structure of
pollen, along with the confirmation of
Giovanni Battista Amici's 1823 discovery of the
pollen-tube, the confirmation of
Robert Brown's views as to the structure of the unimpregnated
ovule (with the introduction of the term "sac embryonnaire", or embryo sac), showing how nearly Brongniart anticipated Amici's subsequent (1846) discovery of the entrance of the pollen-tube into the
micropyle, fertilizing the female cell, which then develops into the
embryo. Of his anatomical works, those of the greatest value are probably the " ("Research on the structure and function of leaves"), and the '
("New research on the epidermis"), in which, among other important observations, the discovery of the cuticle is recorded; and, further, the ' ("Research on the organization of
cycad stems"), giving the results of the first investigation of the anatomy of those plants. His systematic work is represented by a large number of papers and monographs, many of which relate to the
flora of
New Caledonia; and by his '''' (1843), a catalogue of the plants in cultivation at the
French National Museum of Natural History; it is a landmark in the history of classification in that it forms the starting-point of the classification system, modified successively by
Alexander Braun,
August W. Eichler and
Adolf Engler, which was not superseded until the development of DNA research. Brongniart sailed with Louis-Isidore Duperrey aboard the
Coquille, and with Jules Dumont d'Urville and Bory de Saint-Vincent, wrote the botanical atlas for the voyage. In addition to his scientific and professorial labours, Brongniart held various important official posts in connection with the department of education, and interested himself greatly in agricultural and horticultural matters. With
Jean Victoire Audouin and
Jean-Baptiste Dumas, his future brothers-in-law, Brongniart founded the
Annales des Sciences Naturelles, a peer-reviewed journal, in 1824. He also founded the
Société Botanique de France in 1854, and was its first president. Brongniart died in Paris in 1872 and is buried in Division 20 of the
Cimetière du Père Lachaise. ==References==