Agaonidae

The family has changed several times since its taxonomic appearance after the work of Francis Walker in 1846 described from the wasp genus Agaon. Previously the subfamilies Epichrysomallinae, Otitesellinae, Sycoecinae, Sycoryctinae, Sycophaginae, and Agaoninae were the subdivisions of the family. Recent works building strong molecular phylogenies with an extended sampling size have changed the composition of Agaonidae. The paraphyletic groups have been excluded; Epichrysomallinae was raised to family status (Epichrysomallidae), whereas Otitesellinae, Sycoecinae, Sycophaginae, and Sycoryctinae were transferred to Pteromalidae. New subfamilies have been instated (Kradibiinae and Tetrapusiinae). ==Ecology==

The relationship between fig trees and agaonid fig wasps is an obligate mutualism that has evolved over a period of about sixty million years. ==Morphological adaptations==

The pollinating female fig wasps are winged and in general dark, while the males are mostly wingless and whitish. This difference of color is probably due to a clear split in the gender role. Once they have mated, male and female fig wasps have different fates. In some fig species, such as Ficus subpisocarpa or Ficus tinctoria, the males have to chew a hole for the females to leave their natal fig. The winged female wasps can fly over long distances before finding another fig to oviposit in it, while the male dies after chewing a hole. As the fig is closed by a tight ostiole, the female wasps have developed adaptations to enter. First, the mandibles of the female wasps have developed specialized mandibular appendages to help them crawl into the figs. These appendages are adapted to the host fig species, with for instance spiraled ostioles matched by spiral mandibular appendages. ==Subfamilies and genera==

===Agaoninae=== • Agaon • Alfonsiella • Allotriozoon • Blastophaga • Blastophaga psenes (syn. Cynips psenes ) • Courtella • Deilagaon • Dolichoris • Elisabethiella • Eupristina • Eupristina verticillata • Nigeriella • Paragaon • Pegoscapus • Platyscapa • Pleistodontes • Waterstoniella • Wiebesia ===Kradibiinae=== • Ceratosolen • Kradibia (syn. Liporrhopalum ) ===Tetrapusiinae=== • Tetrapus Extinct genera • †Archaeagaon Insect Limestone, United Kingdom, Eocene (Priabonian) • †Archaeagaon minutum (Donisthorpe) == References ==