Agnes Arber

Agnes Robertson was born on 23 February 1879 in Primrose Hill, London. She was the first child of Henry Robert Robertson, an artist, and Agnes Lucy Turner, and had three younger siblings, Donald Struan Robertson (who later became Regius Professor of Greek in the University of Cambridge), Janet Robertson, who later became a portrait painter, and Margaret Robertson (married name Hills), who was a notable suffragist and local politician. Her father gave her regular drawing lessons during her early childhood, which later provided her with the necessary skills to illustrate her scientific publications herself. At the age of eight Robertson began attending the North London Collegiate School founded and run by Frances Buss, one of the leading proponents for girls' education. Under the direction of the school's science teacher Edith Aitken, Robertson discovered a fascination with botany, publishing her first piece of research in 1894 in the school's magazine and later coming first in the school's botany examinations, winning a scholarship. It was here that Robertson first met Ethel Sargant, a plant morphologist who gave regular presentations to the school science club. Sargant would later become her mentor and colleague, having a profound influence on Arber's research interests and methods. In 1897, Robertson began studying at University College, London, gaining her BSc in 1899. After gaining an entrance scholarship Arber became a member of Newnham College, Cambridge and took a further degree in Natural Sciences. She gained first class results in every examination at both universities, along with several prizes and medals from University College, London. After finishing her Cambridge degree in 1902 Robertson worked in the private laboratory of Ethel Sargant for a year, before returning to University College, London as holder of the Quain Studentship in Biology. She was awarded a Doctorate of Science in 1905. Robertson met Edward Alexander Newell Arber (1870–1918) while studying at Newnham College. They married on August 5, 1909, and moved back to Cambridge, where she would remain for the rest of her life. Her only child, Muriel Agnes Arber, was born in 1913, became a geologist, and died in 2004. Arber and her husband had many interests in common, and her marriage was described as 'happy'. Arber was awarded a Research Fellowship from Newnham College in 1912 and published her first book Herbals, their origin and evolution in the same year. Her husband Newall Arber died in 1918 following a period of ill health. Arber never remarried, but continued with her research. She studied in the Balfour Biological Laboratory for Women from her marriage until the laboratory's closure in 1927. Arber maintained a small laboratory in a back room of her house from then until she stopped performing bench research in the 1940s and turned to philosophical study. ==Death==

Agnes Arber died on 22 March 1960, at the age of 81, and is buried in the churchyard of St Andrew's, Girton. ==Scientific career==

Early career Before attending University College, London Arber spent the summer of 1897 working with Ethel Sargant in her private laboratory in Reigate, where Sargant instructed her on microtechniques used to prepare plant specimens for microscopic examination. Arber returned to work in Sargant's laboratory at least once during the summer holidays while she was studying at University College London. Sargant employed Arber between 1902 and 1903 as a research assistant working on seedling structures, during which time in 1903 she published her first paper 'Notes on the anatomy of Macrozamia heteromera' in Proceedings of the Cambridge Philosophical Society. She was an excellent draughtsperson and often illustrated her books herself; she drew about a third of the illustrations in Water Plants. In 1921, the "botanical establishment" denied Arber the 1921 presidency of British association for the advancement of science (BAAS)'s Botany section (Section K). This process was initiated by Frederick Orpen Bower, based on the premise that Arber should not follow Saunders immediately as president as both were women from Cambridge and that there were more senior male botanists up for the presidency. This reflected an antipathy against Arber not only as a female botanist but also as a Cambridge botanist. He claimed that Edinburgh (where the meeting was held) "has the right to expect better than this... To ask Balfour [of Edinburgh] to sit under the Presidency of Mrs. Arber is ridiculous!" Albert Seward met with Arber, who withdrew her candidacy and resigned as the secretary of the BAAS. In 1925 Arber published her third book The Monocotyledons. The Editors of the Cambridge Botanical Handbooks series had asked Ethel Sargant in 1910 to prepare a volume on the monocots for this series. However ill-health and advancing years made it almost impossible for Sargant to complete the book, and in 1918 she suggested Arber to complete the work. The book was preceded by 10 papers in The Annals of Botany detailing the results of her research. Between 1930 and 1942 Arber conducted research into the structure of flowers, where she investigated the structure of many different forms and used morphological information to interpret other flower structures. Her results were published in 10 review papers spanning this period. In 1937 she published a summary of the morphological ideas which had been discussed concerning floral structure, which was considered an important review article for morphological studies. In January 1942 Arber published her last paper involving original botanical research. All of her subsequent publications were entirely concerned with historical and philosophical topics. ==Philosophical studies==

During the Second World War Arber found it difficult to maintain her small laboratory, due to health problems and supplies were becoming more difficult to obtain. Further, she did not believe it was "fair to her neighbours" to have flammable laboratory materials in her home while there was a danger of wartime bombing. This led to her decision to stop performing laboratory work and to concentrate more on philosophical and historical issues. Arber published work on historical botanists, including a comparison between Nehemiah Grew and Marcello Malpighi in 1942, John Ray in 1943 and Sir Joseph Banks in 1945. Arber had been introduced to the work of Goethe while at school and remained fascinated by his ideas about botany. Arber discusses the processes behind forming a concept from research and examines the philosophy of plant morphology. Arber uses this to examine the structure of flowering plants, and proposes the partial-shoot theory of the leaf. According to this theory, each element of the plant is a shoot or a partial shoot. Leaves are partial shoots that show reduced growth capacity. She mentions: "the leaf is a partial-shoot, revealing an inherent urge towards becoming a whole shoot, but never actually attaining this goal, since radial symmetry and the capacity for apical growth suffer inhibition". The parallelism of leaf and shoot dates back to Goethe, who first described compound leaves as in "reality branches, the buds of which cannot develop, since the common stalk is too frail". For Arber, compound leaves are clusters of united partial-shoots. Recent developmental genetic evidence has supported aspects of the partial shoot-theory of the leaf, especially in the case of compound leaves. Her studies on the philosophy of plant morphology led her to take a broader view of the links between science and philosophy. ==Recognition and awards==

In 1921 she accepted the annual position as president of the botany section of the British Association for the Advancement of Science, following Edith Saunders the previous year, but soon resigned after some members indicated that they did not want two successive women presidents. When she was offered the presidency again in 1926, she did not accept. In 1946 she was elected a Fellow of the Royal Society. She was the first woman botanist and third woman overall to receive this honor. In 1948 she was awarded the Gold Medal of the Linnean Society of London. ==Legacy==

In February 1961, Muriel Arber donated some of her mother's surviving materials to the Hunt Institute for Botanical Documentation. A Blue Plaque was installed on her childhood home (9 Elsworthy Terrace, Primrose Hill, London) in 2018. In 2024 one was attached to her Cambridge home where she also carried out much of her research. In May 2024, a new sponsored Agnes Arber PhD thesis prize in comparative biology at the University of Cambridge was created. Professor Sam Brockington, of Cambridge University Botanic Gardens, said that he hoped the prize "will support the next generation of pioneering botanists, following in the footsteps of Agnes". == Selected publications ==

• • • • • • • • • ==See also==