Hesperocyon

Hesperocyon was assigned to Borophagini by Wang et al. in 1999 The Canidae subfamily Hesperocyoninae probably arose out of Hesperocyon to become the first of the three great dogs groups: Hesperocyoninae (~40–30 Ma), Borophaginae (~36–2 Ma), and the Caninae lineage that led to the present-day canids (including grey wolves, foxes, coyotes, jackals and dogs). At least 28 known species of Hesperocyoninae evolved out of Hesperocyon, including those in the following five genera: Ectopocynus (32–19 mya), Osbornodon (32–18 mya), Paraenhydrocyon (20–25 mya), Mesocyon (31–15 mya) and Enhydrocyon (31–15 mya). ==Evolution==

This genus of primitive canids is the ancestor of all later canids. ==Morphology==

This early, canine looked more like a civet or a small raccoon. Its body and tail were long and flexible, while its limbs were weak and short. Still, the build of its ossicles and distribution of its teeth showed it was a canid. It may have been an omnivore—unlike the hypercarnivorous Borophaginae that later split from this canid lineage. Unlike modern canids, Hesperocyon had five fingers and toes and a dew claw deeply set enough to suggest tree climbing capabilities. ==Fossil record==

The oldest fossil evidence was recovered from Saskatchewan dating from 42.5 mya to 31.0 Ma. The youngest fossil was recovered from the Dog Jaw Butte site, Goshen County, Wyoming dating to the Arikareean age (NALMA) of the Oligocene and Miocene 42.5 mya—31.0 Ma. (AEO). == Palaeobiology ==

Palaeopathology A number of individuals of Hesperocyon sp. have been found with evidence of hereditary osteochondroma, which manifested in the individuals as mushroom-like overgrowths of bone in the medial aspect of the distal half of their radii. ==References==