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Alvarezsauroidea

Alvarezsauroidea is a group of small maniraptoran dinosaurs. The group was first formally proposed by Choiniere and colleagues in 2010, to contain the family Alvarezsauridae and non-alvarezsaurid alvarezsauroids, such as Haplocheirus, which is the basalmost of the Alvarezsauroidea. The discovery of Haplocheirus extended the stratigraphic evidence for the group Alvarezsauroidea about 63 million years further in the past. The division of Alvarezsauroidea into the Alvarezsauridae and the non-alvarezsaurid alvarezsauroids is based on differences in their morphology, especially in their hand morphology.

Description
Fossils of alvarezsauroids were described since the 1990s, with many alvarezsauroids being found in Asia and other members also known from North America, South America and possibly Europe. Haplocheirus, for example, was among the largest member of the Alvarezsauroidea. Because of the size of Haplocheirus and its basal phylogenetic position, a pattern of miniaturization for the Alvarezsauroidea is suggested. Miniaturizations are very rare in dinosaurs, but convergently evolved in Paraves. ==Classification==
Classification
The phylogenetic placement of Alvarezsauroidea is still unclear. At first, they were interpreted as a sister group of Avialae (birds) or nested within the group Avialae because they share many morphological characteristics with them, such as a loosely sutured skull, a keeled sternum, fused wrist elements, and a posteriorly directed pubis. This shows that bird-like characteristics were developed multiple times within the Maniraptora. Furthermore, the Alvarezsauroidea had simplified homogenous dentition, convergent with that of some extant insectivorous mammals. More recently, they have been placed within the Coelurosauria basal to the Maniraptora or as a sister taxon of Ornithomimosauria within the Ornithomimiformes. The cladogram below is the latest and most comprehensive analysis of alvarezsaurian evolution, following Meso et al. (2024). {{clade|style=font-size:85%; line-height:75%; |label1=Alvarezsauria |1= ==Geographical distribution==
Geographical distribution
At first, alvarezsauroids were thought to have been originated in South America. However, the discovery of Haplocheirus, and its basal phylogenetic position, as well as its early temporal position, suggests they derived in Asia rather than South America. Xu et al. (2011) suggested that at least three dispersal events of alvarezsauroids took place; one from Asia to Gondwana, one from Gondwana to Asia, and one from Asia to North America. This hypothesis is consistent with faunal interchanges. Their results showed an absence of statistical support for previous biogeographic hypotheses that favour pure vicariance or pure dispersal scenarios as explanations for the distributions of alvarezsauroids across South America, North America and Asia. They instead found that statistically significant biogeographic reconstructions suggest a dominant role for sympatric events ("within area" ones), combined with a mix of vicariance, dispersal and regional extinction. The Asian origin of alvarezsauroids is also bolstered by the discovery of alvarezsaurid specimens from the Turonian-age Bissekty Formation (some of which were named Dzharaonyx in 2022) of Uzbekistan and Bannykus, Tugulusaurus, and Xiyunykus from the Early Cretaceous of China. However, Makovicky et al came to a different conclusion based on the description of a new specimen of Alnashetri, finding support for the group having a Pangaean ancestral distribution shaped by a combination of vicariance and regional extinctions of basal lineages, leading the disjunction distribution of various divergent clades across landmasses. ==References==
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