Pappenheimer was born in
Cedarhurst, New York. He was the eldest son of Dr. Alwin Max Pappenheimer, a distinguished pathologist on the faculty of the College of Physicians and Surgeons at
Columbia University. His brother
John Pappenheimer later became a professor of physiology at
Harvard University, and his sister
Anne Pappenheimer Forbes became an endocrinologist and clinical professor at
Harvard Medical School. Pappenheimer Jr. received his Ph.D. in
organic chemistry from
Harvard University in 1932. He joined Harvard in 1958 as chairman of the Board of Tutors in Biochemical Sciences. == References ==