In 1971,
Walter Coombs concluded that there was only one species of ankylosaurid during the
Campanian age of the Late Cretaceous of North America. He synonymised the species
Anodontosaurus lambei,
Dyoplosaurus acutosquameus, and
Scolosaurus cutleri with
Euoplocephalus tutus. The synonymisation of
Anodontosaurus lambei and
Euoplocephalus tutus was generally accepted and thus CMN 8530 was assigned to
E. tutus. The second study, by Penkalski (2013), named and described
Oohkotokia from
Montana on the basis of remains that were originally thought to be referable to
Euoplocephalus. Penkalski (2013) performed a small
phylogenetic analysis of some
ankylosaurine specimens. The only
Anodontosaurus specimen that was included in this analysis was its holotype.
Anodontosaurus was placed in a
polytomy with the holotype of
Euoplocephalus and some specimens that are referred to it, while
Oohkotokia was placed in a clade with
Dyoplosaurus, and specimens that are thought to represent either
Dyoplosaurus or
Scolosaurus. The DPF specimen, however, was later made the holotype of a new species,
A. inceptus. The following
cladogram is based on a 2015
phylogenetic analysis of the Ankylosaurinae conducted by Arbour and Currie: }} ==See also==