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Bigfin squid

Bigfin squids are a group of rarely seen cephalopods with a distinctive morphology. They are placed in the genus Magnapinna and family Magnapinnidae. Although the family was described only from larval, paralarval, and juvenile specimens, numerous video observations of much larger squid with similar morphology are assumed to be adult specimens of the same family.

Taxonomy
Magnapinna is the sister group to Joubiniteuthis, another little-known deep-sea squid with an unusual body plan and long arms. Both Magnapinna and Joubiniteuthis are monotypic genera within their own families, Magnapinnidae and Joubiniteuthidae, respectively. They are also closely related to the "whip-lash squid" in the families Chiroteuthidae and Mastigoteuthidae.'''' == Physical specimens ==
Physical specimens
The first record of this family comes from a specimen (Magnapinna talismani) caught off the Azores on 10 August 1883. Due to the damaged nature of the find, little information could be discerned, and it was classified as a mastigoteuthid, first as Chiroteuthopsis talismani '', the first known species from the family, with damaged arms During the 1980s, two additional immature specimens were found in the Atlantic (Magnapinna sp. A), and three more were found in the Pacific (Magnapinna pacifica). Researchers Michael Vecchione and Richard Young were the chief investigators of the finds, and they eventually linked them to the two previous specimens, erecting the family Magnapinnidae in 1998, with Magnapinna pacifica as the type species. == Sightings ==
Sightings
The presumed adult stage of Magnapinna is known only from video observations from submersibles, deep sea oil rig cameras, and remotely-operated vehicles (ROVs); no physical specimens have yet been collected, leaving their exact identity unknown. These individuals and the collected juvenile specimens share the large fins and the vermiform arm tips with no suckers, but the iconic elongated arm tips are known only from the presumed adult individuals. Although it has not been directly confirmed whether these squid are the same as the Magnapinna known from specimens, it is largely accepted that they are members of Magnapinnidae. Although observations had been made over a decade earlier, adult bigfin squid only became known to science in 2001, when marine biology student Heather Holston sent footage of what she described as a "21-foot-long squid" to teuthologist Michael Vecchione. The footage had been recorded from an ROV in the Gulf of Mexico in January 2000 at the request of Holston's boyfriend Eric Leveton, who planned on showing it to her. Leveton was a structural engineer aboard the oil-drilling ship Millennium Explorer, who had happened to look into the ROV operation shack when the squid was observed by operators. Although Vecchione initially surmised from Holston's description that the footage might be the first video of a live giant squid (Architeuthis dux), he realized that the video itself portrayed a completely different squid that had no known identity. Further discussions with other cephalopod researchers found no leads on the identity of the squid, and it was thus dubbed the "mystery squid" for a portion of time. Analysis by Vecchione et al of previous footage from submersibles found other video records of bigfin squid, the earliest from 1988. Around the same time, new high-quality footage of a bigfin squid was also recorded off Hawaii by the ROV Tiburon. In December 2001, Vecchione et al published a paper collating these observations; this was also the first paper to identify them as potential members of the Magnapinnidae, which had been named by Vecchione from the juvenile specimens a few years earlier. Independent of Vecchione's publication, Guerra et al published a paper the following year analyzing some of the early bigfin squid footage, and also identified them as potential adult magnapinnids. Viewing close-ups of the body and head, the fins appear to be extremely large, being proportionately nearly as big as those of bigfin squid larvae. While they do appear similar to the larvae, no specimens or samples of the adults have been taken. While their exact identity is unknown, all of the discovered specimens can be observed to have a beige-colored body, translucent fins, near-white tentacles, and dark eyes. Observation timeline The first visual record of an adult bigfin squid was in September 1988. The crew of the submersible Nautile encountered a bigfin squid off the coast of northern Brazil, , at a depth of . In July 1992, the Nautile again encountered these creatures, observing two individuals during a dive off the coast of Ghana at , first at depth and then again at . Both were filmed and photographed. In November 1998, the Japanese crewed submersible Shinkai 6500 filmed another bigfin squid in the Indian Ocean south of Mauritius, at and . Eric Leveton's video, which was later shared with Vecchione, was taken from the remotely operated underwater vehicle (ROV) of the oil-drilling ship Millennium Explorer in January 2000, at Mississippi Canyon in the Gulf of Mexico () at , and allowed for a size estimate. By comparison with the visible parts of the ROV, the squid was estimated to measure with arms fully extended. These were taken in the Pacific Ocean north of Oʻahu, Hawaii (), at . This video and the pre-2001 videos (which had not previously received much scientific attention) were documented by Vecchione et al in a paper that year, and some of the earlier footage was further analyzed by Guerra et al (2002). '' in 2012 Observations of bigfin squid were made in the Great Australian Bight during towed camera and ROV surveys in 2015 and 2017, respectively. In March 2021, during the expedition to document the wreck of the USS Johnston, the submersible DSV Limiting Factor recorded footage of a juvenile bigfin squid from the Philippine Trench at a depth of . This is the deepest observation of any squid and rivalled only by some unidentified cirrate octopods from the same habitat as the deepest observation of any cephalopod. This makes Magnapinna the first squid known to inhabit the hadal zone. The squid was found at a depth of . ROV SuBastian of Schmidt Ocean Institute observed a bigfin squid in close proximity to a black smoker-type hydrothermal vent on 4 April 2023, during the In Search of Hydrothermal Lost Cities expedition. The squid was seen at a depth of . List Date - Dates are given to the level of precision available from the source material - telemetry displayed is used for observations where an ROV video is the primary source Observer - Corresponds either to the person who collected the specimen; the vessel the specimen was collected on, or the vessel that observed the specimen for video-only observations. Location - Given in latitude/longitude when available, otherwise the general region is given. For ROV dives with navigation data available, a link to the corresponding dive track is given, unless a more precise location is specified by a secondary source. Some coordinates from commercial ROVs are specified via UTM rather than latitude/longitude. Depth - Supplied based on precision available from collection method for physical specimens; otherwise, telemetry from ROV observations is used. } || N/A || M. atlantica holotype || || N/A || Captured at Discovery station 13198 #7 || || 2340m || Shinkai 6500 Dive 460 Cruise YK98-08 MODE'98 Leg4 - Atlantis II Fracture Zone - || || 2195m || ROV operating from Millenium Explorer || || 2576m || Gimnaut expedition, RV Atalante supporting || || 1940m || RV Atlantis supporting DSV Alvin, WHOI cruise AT3-58 || || 3380m || RV Western Flyer supporting ROV Tiburon Hawaii Cruise || || 2586m || KAIKO Dive 208 Cruise KR01-12_leg2 || || 3890m || KAIKO Dive 215 Cruise KR01-12_leg2 || || 2750m || Discoverer Deep Seas supporting ROV Hercules 8 || || 3050m || Discoverer Deep Seas supporting ROV Hercules 8 || || N/A || Bergen Museum MAR-ECO cruise super station 46, local station 374 || || RV Atlantis AT15-03 Dive 4195, subadult male || || 1000-2000m || Cruise ANT XXIV-4 || || 2500m || Shell Perdido, Alaminos Canyon Block 857 || || 200-600m || Collected by MOC10 net sampling during M/V Meg Skansi cruise MS8 as part of DEEPEND-RESTORE || || 1961m || NOAA Okeanos Explorer EX2012 Leg3 Dive 13 || || 1578.4m || || Unpublished Serpent Project || 1784m || || Unpublished Serpent Project || 2388m || OCS Study BOEM 2020-022 || || 2278m || OCS Study BOEM 2020-022 || || 4708m || RV Thomas G. Thompson TN309, HADES || || 1687m || Nautilus Expedition NA043 || || 2317m || OCS Study BOEM 2020-022 || || 2178m || RV Investigator || || 2110m || RV Investigator || || 3664m || Pourquoi Pas? || || 1884m || Old Purple Angel Well || || 3060m || REM Etive || || 3002m || REM Etive || || 3056m || REM Etive || || 2294.20m || Gorgon 1 Well || || 100-200m || Pelagic trawl || || 2092m || ROV SuBastian Dive 295, Necker Ridge: Bridge or Barrier || || 3000m || ROV SuBastian Dive 312, The Great Australian Deep-sea Coral and Canyon Adventure || || 6212m || || || 3000m || INDEX 2021, supporting RV Pelagia || || || Dive 10, Windows to the Deep 2:56:25 || || 1931m || ROV SuBastian Dive 501, In Search of Lost Hydrothermal Cities || || 3300m || Tonga Trench Expedition || ==See also==
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