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Loggerhead sea turtle

The loggerhead sea turtle, loggerhead turtle or loggerhead, is a species of sea turtle distributed throughout the world. It is a marine reptile, belonging to the family Cheloniidae. The average loggerhead measures around 90 cm (35 in) in carapace length when fully grown. The adult loggerhead sea turtle weighs approximately 135 kg (298 lb), with the largest specimens weighing about 200 kg (440 lb). The skin ranges from yellow to brown in color, and the shell is typically reddish brown. No external differences in sex are seen until the turtle becomes an adult, the most obvious difference being the adult males have thicker tails and shorter plastrons than the females.

Taxonomy
Carl Linnaeus gave the loggerhead its first binomial name, Testudo caretta, in 1758. Thirty-five other names emerged over the following two centuries, with the combination Caretta caretta first introduced in 1873 by Leonhard Stejneger. The English common name "loggerhead" refers to the animal's large head. The loggerhead sea turtle belongs to the family Cheloniidae, which includes all extant sea turtles except the leatherback sea turtle. The subspecific classification of the loggerhead sea turtle is debated, but most authors consider it a single polymorphic species. Molecular genetics has confirmed hybridization of the loggerhead sea turtle with the Kemp's ridley sea turtle, hawksbill sea turtle, and green sea turtles. The extent of natural hybridization is not yet determined; however, second-generation hybrids have been reported, suggesting some hybrids are fertile. Evolution Although evidence is lacking, modern sea turtles probably descended from a single common ancestor during the Cretaceous period. Like all other sea turtles except the leatherback, loggerheads are members of the ancient family Cheloniidae, and appeared about 40 million years ago. Of the six species of living Cheloniidae, loggerheads are more closely related to the Kemp's ridley sea turtle, olive ridley sea turtle, and the hawksbill turtle than they are to the flatback turtle and the green turtle. Around three million years ago, during the Pliocene epoch, Central America emerged from the sea, effectively cutting off currents between the Atlantic and Indo-Pacific Oceans. The rerouting of ocean currents led to climatic changes as the Earth entered a glacial cycle. Cold water upwelling around the Cape of Good Hope and reduction in water temperature at Cape Horn formed coldwater barriers to migrating turtles. The result was a complete isolation of the Atlantic and Pacific populations of loggerheads. During the most recent ice age, the beaches of southeastern North America were too cold for sea turtle eggs. As the Earth began to warm, loggerheads moved farther north, colonizing the northern beaches. Because of this, turtles nesting between North Carolina and northern Florida represent a different genetic population from those in southern Florida. The distinct populations of loggerheads have unique characteristics and genetic differences. For example, Mediterranean loggerheads are smaller, on average, than Atlantic Ocean loggerheads. North Atlantic and Mediterranean loggerhead sea turtles are descendants of colonizing loggerheads from Tongaland, South Africa. South African loggerhead genes are still present in these populations today. ==Description==
Description
The loggerhead sea turtle is the world's largest hard-shelled turtle, slightly larger at average and maximum mature weights than the green sea turtle and the Galapagos tortoise. It is also the world's second largest extant turtle after the leatherback sea turtle. Adults have an approximate weight range of , averaging around , and a straight-line carapace length range of . The skull is most easily distinguished from other sea turtles by having maxillae that meet in the mid-line of the palate. The portion of skull behind the eyes is also relatively large and bulbous due to the extensive jaw muscles. ==Distribution==
Distribution
The loggerhead sea turtle has a cosmopolitan distribution, nesting over the broadest geographical range of any sea turtle. It inhabits the Atlantic, Indian, and Pacific Oceans and the Mediterranean Sea. Pacific loggerheads occasionally nest in Vanuatu and Tokelau. Yakushima Island is the most important site, with three nesting grounds visited by 40% of all nearby loggerheads. Greece is the most popular nesting site along the Mediterranean, with more than 3,000 nests per year. Because of this, Greek authorities do not allow planes to take off or land at night in Zakynthos due to the nesting turtles. Another records one specimen being washed up on a beach in County Donegal, Ireland, in 2019. They are not native to UK waters, but have been increasingly seen, most recently in 2026. ==Habitat==
Habitat
Loggerhead sea turtles spend most of their lives in the open ocean and in shallow coastal waters. They rarely come ashore besides the females' brief visits to construct nests and deposit eggs. Hatchling loggerhead turtles live in floating mats of Sargassum algae. Adults and juveniles live along the continental shelf as well as in shallow coastal estuaries. In the northwestern Atlantic Ocean, age plays a factor in habitat preference. Juveniles are more frequently found in shallow estuarine habitats with limited ocean access compared to non-nesting adults. Loggerheads occupy waters with surface temperatures ranging from during non-nesting season. Temperatures from are most suitable for nesting females. Juvenile loggerheads share the Sargassum habitat with a variety of other organisms. The mats of Sargassum contain as many as 100 different species of animals on which the juveniles feed. Prey found in Sargassum mats may include barnacles, crab larvae, fish eggs, and hydrozoan colonies. Some prey, such as ants, flies, aphids, leafhoppers, and beetles, are carried by the wind to the mats. Marine mammals and commercial fishes, including tuna and mahi-mahi, also inhabit the Sargassum mats. ==Behavior==
Behavior
Loggerhead sea turtles observed in captivity and in the wild are most active during the day. In captivity, the loggerheads' daily activities are divided between swimming and resting on the bottom. While resting, they spread their forelimbs to about midstroke swimming position. They remain motionless with eyes open or half-shut and are easily alerted during this state. At night, captives sleep in the same position with their eyes tightly shut, and are slow to react. Female-female aggression, which is fairly rare in other marine vertebrates, is common among loggerheads. Ritualized aggression escalates from passive threat displays to combat. This conflict primarily occurs over access to feeding grounds. Escalation typically follows four steps. It has a greater list of known prey than any other sea turtle. Other food items include sponges, corals, sea pens, polychaete worms, tube worms, sea anemones, cephalopods, barnacles, brachiopods, amphipods, isopods, Portuguese men o' war, insects, bryozoans, hydrozoans, sea urchins, sand dollars, sea cucumbers, starfish, tunicates, fish (eggs, juveniles, and adults), hatchling turtles (including members of its own species), algae, and vascular plants. During migration through the open sea, loggerheads eat jellyfish, floating molluscs, floating egg clusters, squid, and flying fish. is a predator of loggerhead nests in Australia. Loggerheads have numerous predators, especially early in their lives. Egg and nestling predators include ghost crabs, oligochaete worms, some beetles, flesh fly larvae, some ants, flesh flies, snakes, gulls, corvids, opossums, bears, rats, armadillos, mustelids, skunks, canids like coyotes, dingos, the Red foxes in Australia, Jackals and feral dogs, procyonids, Feral cats, Feral pigs, and humans. During their migration from their nests to the sea, hatchlings are preyed on by dipteran larvae, crabs, toads, lizards, snakes, seabirds such as frigatebirds, and other assorted birds and mammals. In the ocean, predators of the loggerhead juveniles include portunid crabs and various fishes, such as parrotfishes and moray eels. Adults are more rarely attacked due to their large size, but may be preyed on by large sharks (such as bull sharks, oceanic whitetip sharks, tiger sharks, and great white sharks), monk seals, and killer whales. Nesting females are attacked by flesh flies, feral dogs, and humans. Salt marsh mosquitos can also pester nesting females. ==Life history==
Life history
Early life Hatchlings range in color from light brown to almost black, lacking the adult's distinct yellows and reds. Like all marine turtles, the loggerhead prepares for reproduction in its foraging area. This takes place several years before the loggerhead migrates to a mating area. Female loggerheads first reproduce at ages 28–33 in Southeastern United States and Australia, and at ages 17–30 in South Africa. Age at first reproduction in the Mediterranean, Oman, Japan, and Brazil are unknown. Nesting loggerheads have a straight carapace length of . Because of the large range, carapace length is not a reliable indicator of sexual maturity. Their estimated maximum lifespan is 47–67 years in the wild. Reproduction Female loggerheads first reproduce between the ages of 17 and 33, and their mating period may last more than six weeks. They court their mates, but these behaviors have not been thoroughly examined. Male forms of courtship behavior include nuzzling, biting, and head and flipper movements. Studies suggest females produce cloacal pheromones to indicate reproductive ability. Before mating, the male approaches a female and attempts to mount her, while she resists. Next, the male and female begin to circle each other. If the male has competitors, the female may let the males struggle with each other. The winner then mounts the female; the male's curved claws usually damage the shoulders of the female's shell during this process. Other courting males bite the male while he is attempting to copulate, damaging his flippers and tail, possibly exposing bones. Such damage can cause the male to dismount and may require weeks to heal. While nesting, females produce an average of 3.9 egg clutches, and then become quiescent, producing no eggs for two to three years. Unlike other sea turtles, courtship and mating usually do not take place near the nesting beach, but rather along migration routes between feeding and breeding grounds. Recent evidence indicates ovulation in loggerheads is mating-induced. Through the act of mating, the female ovulates eggs which are fertilized by the male. This is unique, as mating-induced ovulation is rare outside of mammals. In the Northern Hemisphere, loggerheads mate from late March to early June. The nesting season is short, between May and August in the Northern Hemisphere and between October and March in the Southern Hemisphere. Loggerheads may display multiple paternity. Multiple paternity is possible due to sperm storage. The female can store sperm from multiple males in her oviducts until ovulation. A single clutch may have as many as seven fathers, each contributing sperm to a portion of the clutch. Multiple paternity and female size are positively correlated. Two hypotheses explain this correlation. One posits that males favor large females because of their perceived higher fecundity (ability to reproduce). The other states, because larger females are able to swim more quickly to mating grounds, they have longer mating periods. All sea turtles have similar basic nesting behaviors. Females return to lay eggs at intervals of 12–17 days during the nesting season, on or near the beach where they hatched. They exit the water, climb the beach, and scrape away the surface sand to form a body pit. With their hind limbs, they excavate an egg chamber in which the eggs are deposited. The females then cover the egg chamber and body pit with sand, and finally return to the sea. This process takes one to two hours, and occurs in open sand areas or on top of sand dunes, preferably near dune grasses that the females can use to camouflage the nest. The nesting area must be selected carefully because it affects characteristics such as fitness, emergence ratio, and vulnerability to nest predators. Loggerheads have an average clutch size of 112.4 eggs. ==Conservation==
Conservation
Many human activities have negative effects on loggerhead sea turtle populations. The prolonged time required for loggerheads to reach sexual maturity and the high mortality rates of eggs and young turtles from natural phenomena compound the problems of population reduction as a consequence of human activities. This raises concern over the connection between rapid global temperature changes and the possibility of population extinction. The increase of temperature and food availability will increase reproduction output of loggerhead turtles. Many researchers agree that temperature increases due to climate change has a complicated impact on turtles. At breeding sites when a loggerhead turtle lays multiple clutches in a season, a higher temperature will cause the duration of time between laying two different nests to become shorter. The amount of food availability makes a difference in reproductive output because when there is a greater amount of food available, the turtles will grow to a larger size. The larger a turtle is, the more likely they will have a greater reproductive output. The amount of food also has a relationship to temperature. Researchers have found that an increase of temperature causes feeding grounds to produce more food. Tropical Cyclones have a significant impact on hatchling loss. The associated storm surges push water higher up the beach, flooding nest and drowning the embryos. Strong wave action may eroded away sand, exposing the eggs to drying and predation. The current trend of rising sea surface temperatures and the increase in both numbers and intensities of tropical cyclones as a result of climate change pose a growing threat to turtle populations. Using ONT nanopore direct-DNA sequencing the blood methylome profile was also derived. The researchers finding the microchromosomes are particularly useful for monitoring functional genetic and epigenetic diversity. United States The National Marine Fisheries Service (NMFS), National Oceanic and Atmospheric Administration (NOAA), the U.S. Fish and Wildlife Services (USFWS), and the Department of the Interior ruled four distinct population segments as threatened (Northwestern Atlantic Ocean, South Atlantic Ocean, Southeast Indo-Pacific Ocean, and Southwest Indian Ocean) and five as endangered (Mediterranean Sea, North Indian Ocean, North Pacific Ocean, Northeast Atlantic Ocean, and South Pacific Ocean) effective on October 24, 2011. Off the coast of southern California NMFS, NOAA, and Department of Commerce prohibited fishing with large drift gillnet (DGN) gear in the loggerhead conservation area during the presence of El Niño conditions in order to protect the endangered North Pacific Ocean loggerhead DPS. This ruling effective July 23, 2014 was intended to prevent bycatch of loggerhead sea turtles. Critical habitat designation for the Northwest Atlantic Ocean DPS of loggerhead sea turtles specified 38 marine areas that include nearshore reproductive habitat, breeding areas, winter area, constricted migratory corridors, and Sargassum habitat. This ruling was made the NMFS, NOAA, and Department of Commerce effective August 11, 2014. The 2012 BiOp is an integral component to managing the shallow-set fishery, because the one-year incidental take statement (ITS, including reasonable and prudent management measures, and terms and conditions) forms the basis for regulations that specify the annual limits on leatherback and North Pacific loggerhead sea turtle interactions with the fishery that are necessary to manage the impacts of the fishery on sea turtles. Effective January 11, 2010 the NMFS, NOAA, and Department of Commerce removed the limit on the number of fishing gear deployments for the Hawaii-based pelagic shallow-set longline fisheries and simultaneously increased the number of incidental interactions allowed with loggerhead sea turtles. This ruling stated that longline fisheries may not interact with over 46 loggerhead sea turtles a year, a number thought to not interfere with survival and recovery of loggerhead sea turtles. Incidental interaction limit for loggerhead turtles was increased from 17 to 34 interactions a year starting November 5, 2012. ==Symbols==
Symbols
The loggerhead sea turtle appears on the $1000 Colombian peso coin. In the United States, the loggerhead sea turtle is the official state reptile of South Carolina and also the state saltwater reptile of Florida. ==See also==
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