in the oceans surrounding Southern Africa. Note the very high concentrations along the west coast, due to the upwelling of nutrient-rich water from the cold depths of the South Atlantic Ocean, forming the Benguela Current.Northward winds along the coast result in
Ekman transport offshore and
upwelling of nutrient-rich deep water to the
euphotic zone. The intensity of the upwelling event is determined by wind strength. Variations in wind strength result in pulses of upwelling, which propagate to the south along the coast with speeds of 5 to 8 m/s. The pulses are similar to a
Kelvin wave, except on a scale of 30 to 60 km instead of 1000 km, and can propagate around the Cape depending on wind systems. Pulses of upwelling induce biological production. In the Benguela system,
phytoplankton growth requires a period of upwelling followed by a period of
stratification and relatively calm waters. The
phytoplankton bloom usually lags the upwelling event by 1 to 4 days and blooms for 4 to 10 days. In order for
zooplankton to have a continuous food supply, the phytoplankton blooms must not occur too far apart. Pulses of upwelling in the Benguela system regularly have a duration of 10 days, an optimal period for biological production. It is estimated that the annual
biomass production in the Benguela system is 4.7×
gC/y, making the Benguela system 30 to 65 times more productive per unit area than the global ocean average. While upwelling promotes abundant
primary and
secondary production in the upper parts of the water column and near the coast, deeper waters with limited oxygen exchange create hypoxic areas called
oxygen minimum zones at the coastal shelf and upper coastal slope. The Benguela oxygen minimum zone starts around a depth of 100 m and is a few hundred meters thick. Bacteria that use sulphur rather than oxygen, called
sulphur-reducing bacteria, reside in the oxygen minimum zone. The most abundant fish in the Benguela system are
Sardinops and
Engraulis. The
Southern African pilchard (
S. s. ocellatus), was intensely fished in the 1950s and peaked in 1968 with landings over 1.3 million tons. Since then, fishery of the
Sardinops has declined and fishery of the
Southern African anchovy (
Engraulis capensis) has taken over. ==Benguela Niño==