Most of the main cast from the show's
third season returned for the fourth season, with the exception of
Ashley Jensen.
Daniel Eric Gold was promoted to the main cast as Matt Hartley, having appeared in a recurring capacity during Season 3.
Yaya DaCosta took over the recurring role of Nico Slater, who was played in the first season by
Jowharah Jones. The fourth season saw guest appearances by the brothers of two of the show's stars. Becki Newton's brother Matt Newton took on the recurring role of Marc's new boyfriend Troy, and Vanessa Williams's brother
Chris Williams played drag queen "Wilheldiva Hater".
Main Cast •
America Ferrera as
Betty Suarez •
Vanessa L. Williams as
Wilhelmina Slater •
Eric Mabius as
Daniel Meade •
Judith Light as
Claire Meade •
Michael Urie as
Marc St. James •
Becki Newton as
Amanda Tanen •
Ana Ortiz as
Hilda Suarez •
Tony Plana as
Ignacio Suarez •
Mark Indelicato as
Justin Suarez •
Daniel Eric Gold as Matt Hartley (episodes 1-11)
Recurring Cast •
Dylan Baker as Bennett Wallis •
Neal Bledsoe as Tyler Venton •
Grant Bowler as Connor Owens •
Smith Cho as Megan •
Yaya DaCosta as Nico Slater •
Kristen Johnston as Helen •
Ralph Macchio as Councilman Archie Rodriguez •
Alec Mapa as Suzuki St. Pierre • Piter Marek as Jonathan Vieja •
Ryan McGinnis as Austin Marley •
Matt Newton as Troy •
David Rasche as Calvin Hartley •
Adam Rodriguez as Bobby Talercio •
Jamie-Lynn Sigler as Natalie
Guest Cast •
Christie Brinkley as Penelope Graybridge •
Brooklyn Decker as Lexie •
Christine Ebersole as Frances •
Tovah Feldshuh as Mrs. Varner •
Jesse Tyler Ferguson as Dr. Gabe Farkus •
Christopher Gorham as
Henry Grubstick •
Dana Ivey as Roberta •
Ashley Jensen as
Christina McKinney •
Carol Kane as Lena Korvinka •
Lainie Kazan as Dina Talercio •
Sarah Lafleur as Molly Meade •
Hamish Linklater as Evan Grant •
Brian Stokes Mitchell as Donald Jones •
Donna Murphy as Eve •
Kathy Najimy as Dr. Frankel •
Lynn Redgrave as Olivia Guillemette •
Freddy Rodriguez as Giovanni ‘Gio’ Rossi •
RuPaul as Rudolph •
Rich Sommer as Jimmy Wilson •
Fisher Stevens as Mr. Z •
Aaron Tveit as Zachary Boule •
Lauren Vélez as Elena Sanchez •
Chris Williams as Wilheldiva Hater In addition, the final season featured cameo appearances by singer
Shakira, designer Brian Reyes, and drag queens
Varla Jean Merman, Candi Shell, and
Hedda Lettuce as themselves. ==Episodes==