The holotype of
Brachypterygius extremus is a single right forepaddle, clearly different from other Late Jurassic ichthyosaurs (e.g.
Ophthalmosaurus).
Species previously assigned to Brachypterygius Owen (1840) erected
Ichthyosaurus trigonus based upon a single dorsal vertebra (
ANSP 10124) from the Kimmeridge Clay of Westbrook,
Wiltshire, UK. The holotype was long thought to be lost until it was rediscovered in 1988. Many specimens were referred to
Ichthyosaurus trigonus; Bauer (1898) suggested that – what are now –
Ophthalmosaurus,
Brachypterygius and
Nannopterygius should be synonymised into
Ichthyosaurus trigonus.
I. trigonus was included in the new genus
Macropterygius by Huene (1922), who later (1923) made it the type species of the genus. Nowadays,
I. trigonus (and hence
Macropterygius) is a nomen dubium because its holotype is indistinguishable from other ophthalmosaurids. (Christopher McGowan and Ryosuke Motani mistakenly stated that
I. trigonus may be synonymous with
Ophthalmosaurus icenicus McGowan and Motani (2003) considered it to be a species of
Brachypterygius, In 2020, Nikolay Zverkov and Dmitry Grigoriev reassigned this species to
Maiaspondylus. A large skull was discovered in the Kimmeridge Clay of
Stowbridge,
Norfolk, UK and named as a new genus and species,
Grendelius mordax, by McGowan in 1976. ==Phylogeny==