during testing in 2011 During early 2011, the Class 379 began its phased introduction into revenue service. On 17 March 2011, the first two units, 379005 and 379006, performed the type's first service with a high-profile launch by Secretary of State for Transport
Philip Hammond. 379005 was named
Stansted Express in a ceremony at . Over the following three months, a further 18 sets joined the initial pair working the Stansted Express services. They have been worked in a mixture of 12-car and 8-car lengths. The fleet was maintained by Bombardier at
Ilford EMU Depot. The new fleet's introduction was delayed: the first Class 745/1 entered service on 28 July 2020. Due to high leasing costs Greater Anglia withdrew the Class 379 fleet from service in February 2022, after which the units were placed in storage. In March 2024,
Porterbrook announced that they had purchased the units from Akiem, and that
Govia Thameslink Railway planned to use the trains on
Great Northern services. In November 2024, Great Northern announced that all Class 379s had been accepted into their fleet. The trains entered service on Great Northern route services on 10 February 2025.
Battery-electric multiple unit trial During 2013, the national infrastructure owning company
Network Rail announced that unit 379013 would be used as a testbed for a future
battery electric multiple unit. Following several months of conversion work and non-service testing, the unit was used to carry passengers for the first time on a
Manningtree–Harwich Town service on 12 January 2015. Throughout its five-week trial period, data was gathered to assess its performance; it could reportedly operate for up to an hour on battery power alone, while charging via the pantograph took two hours. ==Fleet details==