Placement within Bovini }} }} The majority of phylogenetic work based on
ribosomal DNA,
chromosomal analysis, autosomal
introns and
mitochondrial DNA has recovered three distinctive subtribes of
Bovini: Pseudorygina (represented solely by the
saola), Bubalina, and Bovina (which today are represented by the genera
Bison and
Bos). One
cytogenetic analysis concerning the phylogenetic position on the saola suggests the species could be related to buffalo. This relationship has not, however, been supported by most phylogenetic work concerning Bovini. since 2011 new uncertainty over the number of species that should be recognized has been introduced. with the African buffalo split into five subspecies based on differences in the horns and skin colouration. The bovid biologist Castelló adopted the Groves and Grubb taxonomic interpretation, Below is the listing of 'new' species recognized by Groves and Grubb (2011) Below is the list of a number of the described fossil species (listed alphabetically): •
Subtribe Bubalina (Rütimeyer, 1865) :* Genus
Bubalus (Smith, 1827) ::* †
Bubalus brevicornis (Young, 1936) ::* †
Bubalus cebuensis (Croft
et al., 2006) ::* †
Bubalus grovesi (Rozzi, 2017) ::* †
Bubalus mephistopheles (Hopwood, 1925) ::* †
Bubalus murrensis (Berckhemer, 1927) European buffalo, extinct c. 11,000 years Before Present ::* †
Bubalus palaeindicus (Falconer, 1859) ::* †
Bubalus palaeokerabau (Dubois, 1908) ::* †
Bubalus platyceros (Lydekker, 1877) ::* †
Bubalus teilhardi (Young, 1932) ::* †
Bubalus wansijocki (Boule & Teilhard, 1928) ::* †
Bubalus youngi (Chow & Hsu, 1957) :* Genus †
Hemibos (Falconer, 1865) ::* †
Hemibos acuticornis (Falconer & Gautley, 1868) ::* †
Hemibos antelopinus (Falconer & Gautley, 1868) ::* †
Hemibos galerianus (Petronio & Sardella, 1998) ::* †
Hemibos gracilis (Qiu, 2004) ::* †
Hemibos triquetricornis (Falconer, 1865) :* Genus †
Parabos (Arambourg & Piveteau, 1929) ::* †
Parabos cordieri (de Christol, 1832) ::* †
Parabos macedoniae (Arambourg & Piveteau, 1929) ::* †
Parabos soriae (Morales, 1984) :* Genus †
Proamphibos (Pilgrim, 1939) ::* †
Proamphibos hasticornis (Pilgrim, 1939) ::* †
Proamphibos kashmiricus (Pilgrim, 1939) ::* †
Proamphibos lachrymans (Pilgrim, 1939) :* Genus
Syncerus (Hodgson, 1847) ::* †
Syncerus acoelotus (Gentry & Gentry, 1978) ::* †
Syncerus antiquus (Duvernoy, 1851) – formerly a species of
Pelorovis :* Genus †
Ugandax (Cooke & Coryndon, 1970) ::* †
Ugandax coryndonae (Gentry, 2006) ::* †
Ugandax demissum (Gentry, 1980) ::* †
Ugandax gautieri (Cooke & Coryndon, 1970) ==Feral buffaloes==