MarketCarcinosoma
Company Profile

Carcinosoma

Carcinosoma is a genus of eurypterid, an extinct group of aquatic arthropods. Fossils of Carcinosoma are likely restricted to deposits of late Silurian to early Devonian age, although an Early Ordovician specimen has been classified as ?Carcinosoma aurorae, which would make this the earliest known eurypterid genus. Classified as part of the family Carcinosomatidae, which the genus lends its name to, Carcinosoma contains several species from North America and Great Britain.

Description
Carcinosoma was among the largest eurypterids, with isolated fossil remains consisting of a long metastoma (a plate overlaying the coxae of the first six appendages) of the species C. punctatum indicating a full length of . Fossil prosomal appendages (appendages attached to the head) referred to the species could possibly increase this estimate to an overall length of around . Other species of Carcinosoma were smaller, most being in the range of to in length. Indeed, the walking legs (the second to fifth pair of appendages) were stout and strong and increased in size anteriorly, from the fifth to third pair of appendages, though the first pair of appendages were much shorter than the following pairs. As such, the second pair of walking legs were the longest. Each walking appendage possessed long and curved spines, often two such spines occurring per joint.'''|alt= Carcinosoma ranged in size from to in length, with the largest species by far being C. punctatum, known from the Ludlow epoch of England, and the smallest being C. libertyi, known from the Late Llandovery epoch of Canada. Though no other species came close to the size of C. punctatum, many species were moderately large, including C. harleyi from the Late Ludlow epoch of England at , and C. newlini from the Early Pridoli epoch of the United States at . == History of research ==
History of research
parts of the telson) Carcinosoma was first described under the name Eurysoma (meaning "wide body", deriving from Greek εὐρύς, "wide", and Latin soma, "body") by British American geologist and paleobotanist Edward Waller Claypole in 1890, who named the type species of the new genus E. newlini in honor of a C. E. Newlin who had collected the fossils. The Eurysoma specimens had been discovered in deposits of Early Pridoli age in the Kokomo Formation of Indiana alongside several other eurypterid specimens, all of which at the time were referred to Eurypterus lacustris (though Claypole noted in the same paper that this may have been done hastily). Later in the same year, Claypole discovered that the name Eurysoma was preoccupied and thus not available to be used for his genus of eurypterids. Claypole replaced the name Eurysoma with the new name Carcinosoma. and soma, "body". Another species of Eurypterus, E. scoticus was named in 1899 by Scottish zoologist and paleontologist Malcolm Laurie based on fragmentary remains recovered in deposits of Llandovery age in Scotland. In 1912, American paleontologists John Mason Clarke and Rudolf Ruedemann noted that Carcinosoma was sufficiently similar to the related eurypterid Eusarcus to be designated as synonymous with it. As Eusarcus had been named in 1875, fifteen years earlier than Carcinosoma, its name had priority and replaced Carcinosoma. At this time, the combined genus of Eusarcus contained several species that are today seen as Carcinosoma, including C. newlini, C. scoticus and C. scorpioides, which Clarke and Ruedemann had referred to the genus on account of their similarities with C. newlini and species previously referred to Eusarcus. In 1934, 59 years after it had been described, Eusarcus was recognized as a name preoccupied by a harvestman. The Norwegian geologist Leif Størmer proposed that the name of the taxon should be next oldest available and valid name for the genus, Carcinosoma. During the preparation for his paper on the issue, Størmer also discussed the situation with fellow Norwegian researcher Embrik Strand, who helped confirm that Carcinosoma was not preoccupied. Strand would subsequently propose the replacement name Eusarcana in 1942, despite the problem having been dealt with by Størmer, who he had been in contact with, eight years earlier. The reasons for proposing the name during the circumstances of the time remains unknown, but critique from contemporary researchers of Strand for his studies in systematics and an apparent desire to name as many taxa as possible may explain the situation somewhat. As it was seen as completely unnecessary at the time, Strand's Eusarcana was overlooked and not even mentioned in subsequent eurypterid studies. In 1961, American paleontologist Erik N. Kjellesvig-Waering classified Eurypterus punctatus (originally described as Pterygotus punctatus by English paleontologist and prominent eurypterid researcher John William Salter in 1859) as Carcinosoma punctatum and named a new species C. harleyi based on fossils from the Ludlow epoch of the Welsh Borderland. Kjellesvig-Waering could differentiate C. harleyi from C. punctatum based on C. harleyi lacking serrations on the eighth podomere of the swimming leg and the serrations of the ninth podomere being less developed. C. punctatum was diagnosed by Kjellesvig-Waering in 1961 based on the considerably pronounced serrations of the distal parts of its swimming leg, but the diagnosis is only valid for the lectotype specimen of the species, BGS GSM89435 (compromising the distal parts of a swimming leg discovered in deposits of Middle Ludlow age in the Mocktree Shale of Leintwardine in Herefordshire, England), and four other specimens from the same locality (BMNH 39389, BMNH In. 43804, BGS GSM89561 and GSM89568). Due to a close resemblance of the swimming legs, C. punctatum is assumed to have been similar in appearance to C. newlini. C. punctatum can be distinguished from C. newlini by the serration along the margin of the distal podomeres of C. punctatum being more pronounced. C. harleyi, from the Late Ludlow epoch, was described mainly based on specimens previously known (some having been reported by Salter as early as 1859) but previously referred to Eurypterus punctatus. Recognized by Kjellesvig-Waering as distinct, the species is named in honor of John Harley, one of the earliest collectors of eurypterid fossils in the region. Noted as moderately large in size by Kjellesvig-Waering, the holotype specimen of C. harleyi (No. 89434 in the collection of the Geological Survey and Museum in London) is a fragment of a swimming leg measuring in length. The nearly complete lack of serrations in the joints of C. harleyi makes the species very distinct from C. punctatum and other species of Carcinosoma. In 1964, both C. punctatum and C. harleyi were still recognized as part of Carcinosoma following an emended diagnosis of the genus by Kjellesvig-Waering and American paleontologist Kenneth Edward Caster, though C. harleyi was only tentatively recognized. Further species recognized at the time were C. libertyi and C. logani (both from Ontario, Canada; C. logani was later found to be a crustacean and not a eurypterid at all), C. spiniferum (from New York, United States), C. newlini (from Indiana, United States), C. scorpioides and C. scoticus (both from Scotland). Out of these species, only C. newlini and C. scorpioides preserve the swimming legs, where the diagnostic characters of the genus are, which makes the assignment of the other species to Carcinosoma less secure. Kjellesvig-Waering and Caster also recognized Eusarcus and Carcinosoma to be distinct genera when revising the superfamily Carcinosomatoidea, and coined the replacement name Paracarcinosoma to designate the species previously assigned to Eusarcus. E. scorpionis was designated the type species. Caster and Kjellesvig-Waering made no mention of Embrik Strand or Eusarcana, and they were likely not aware of the existence of the previous name. In 2012, American paleontologists Jason A. Dunlop and James Lamsdell designated Paracarcinosoma as a junior synonym of Eusarcana per the taxonomic laws of priority. In 2025, Lamsdell reclassified C. spiniferum as a species of Eusarcana and reclassified both C. scorpioides and C. scoticus as species of the new genus Cruinnopterus.'''''' In 2025, Belgian paleontologist Peter Van Roy, along with his coauthors Jared C. Richards and Javier Ortega-Hernández described isolated appendages of a carcinosomatid eurypterid from the Early Ordovician Fezouata biota of Morocco. They tentatively assigned these specimens to Carcinosoma due to their robustness and erected the new species ?Carcinosoma aurorae, diagnosed by the paired articulating spines being of unequal length, the "positions of long and short spines alternating between podomeres," and the teeth of the gnathobases being "long and slender," to contain them. == Classification ==
Classification
Carcinosoma is classified as part of the family Carcinosomatidae, a family within the superfamily Carcinosomatoidea, alongside the genera Eusarcana, Eocarcinosoma, Rhinocarcinosoma and possibly Holmipterus. The cladogram below is adapted from a larger cladogram (simplified to only display the Carcinosomatoidea) in a 2007 study by eurypterid researcher O. Erik Tetlie, in turn based on results from various phylogenetic analyses on eurypterids conducted between 2004 and 2007. The second cladogram below is simplified from a study by Lamsdell et al. (2015). Tetlie (2007) 'Lamsdell et al. (2015)' {{clade| == Paleobiology ==
Paleobiology
of C. newlini and a close-up of the "post-telson" with its segments labelled 1-11 The walking legs of Carcinosoma were turned forward, which also directed the large spines on the appendages forward. In C. newlini, these flat and forward-facing legs are thought to have been used to create a trap to capture prey in. The strong structures seen in C. newlini are not reflected in other carcinosomatids. For instance, the appendages of Eusarcana were much more weakly developed and would not have served as an effective weapon. Eusarcana is more likely to have relied on its telson, taking the shape of a sharp and curved stinger similar to that of scorpions and potentially capable of injecting venom. == Paleoecology ==
Paleoecology
(head) and walking appendages of a fossil specimen of C. newlini As the opisthosoma of Carcinosoma wasn't as streamlined as that of more active eurypterids and on account of its unique telson morphology, it is believed that Carcinosoma was not a very active swimmer. It is unlikely to have been well adapted to a completely nektonic (actively swimming) lifestyle and is more likely to have been nektobenthic (swimming near the bottom). Complex eurypterid faunas, compromising several different species in different ecological roles, are typical of the period. These faunas were typically dominated by one or more particular eurypterid families, the dominant groups depending on the environment and location. Three such types of eurypterid faunas have been documented from the late Silurian, out of which a Carcinosomatidae-Pterygotidae fauna is the most marine type. All known examples of Carcinosoma are known from marine beds, typically occurring with trilobites, starfish, bryozoans, brachiopods, linguloids and other marine animals. Carcinosoma also prominently occurs together with pterygotid eurypterids. In the fossil deposits of the Welsh Borderland, examples of Carcinosoma occur together with representatives of the pterygotid genera Erettopterus and Pterygotus over a period of millions of years (though other eurypterids, such as Salteropterus, Dolichopterus, Hughmilleria, Eurypterus, Marsupipterus, Mixopterus, Parahughmilleria, Slimonia, Tarsopterella and Stylonurus are also present in lesser numbers). Other types of late Silurian eurypterid faunas include one dominated by the Eurypteridae. When genera such as Erieopterus or Eurypterus occur in great numbers other genera and families are more rare, though groups such as dolichopterids, carcinosomatids and pterygotids tend to occur in small numbers. The environments with such faunas appear to be quieter waters such as lagoons, estuaries and bays. The third and final recognized type of fauna is one dominated by hughmilleriids and stylonurids, generally alongside sandy bottoms and with few other associated fossils. The environments inhabited by this third fauna was likely less marine than the others, possibly representing the more brackish parts of bays and estuaries. == See also ==
tickerdossier.comtickerdossier.substack.com