Sphenophyllales are small, slender branching plants, usually growing to a height of less than tall. The long stems range from to in diameter. The stems are jointed and ribbed with weak
habits, making it probable that these plants were
vine or shrub-like when alive, and formed a portion of the understory in Carboniferous forests. The stem anatomy is
protostelic (
root-like), containing a solid primary
xylem core with secondary xylem tissue present in some species. The leaves, which can be several centimeters long, are borne on each node in whorls (called verticels) and are wedge-shaped, fan-shaped, linear, or forked. Reproductive parts are either long terminal cones (consisting of two lobes, a sterile lower lobe and the fertile upper lobe bearing the
sporangia) or loose
strobili. All sphenophylls are
homosporous, with monolete or trilete
spores.{{cite book ==Taxonomy==