chewing cud The
alimentary canal of ruminants, such as
cattle,
giraffes,
goats,
sheep,
alpacas, and
antelope, are unable to produce the
enzymes required to break down the
cellulose and
hemicellulose of plant matter. Accordingly, these animals rely on a
symbiotic relationship with a wide range of
microbes, which largely reside in the
reticulorumen, and which can synthesize the requisite enzymes. The reticulorumen thus hosts a microbial fermentation which yields products (mainly
volatile fatty acids and microbial protein), which the ruminant is able to digest and absorb. This allows the animals to extract nutritional value from cellulose which is usually undigested. The process of rumination is stimulated by the presence of
roughage in the upper part of the
reticulorumen. The chest cavity is stretched, forming a vacuum in the gullet that sucks the semi-liquid stomach content into the
esophagus. From the esophagus it is taken back to the mouth with retro
peristaltic movements. When the stomach content, or the cud, arrives in the mouth of the ruminant, it is pushed against the palate with the tongue to remove excess liquid, the latter is swallowed and the solid material is chewed thoroughly so the cattle can extract the minerals present in the cud brought to the surface during rumination. The function of rumination is that food is physically refined to expose more surface area for bacteria working in the reticulorumen, as well as stimulation of
saliva secretion to
buffer the rumen
pH. When food has been degraded sufficiently it passes from the reticulorumen through the reticulomasal orifice to the
omasum followed by the
abomasum to continue the digestion process in the lower parts of the alimentary canal. No enzymes are secreted in the rumen. Enzymes and
hydrochloric acid are only secreted from the
abomasum (fourth stomach) onwards, and ruminants function from that point onwards much like monogastric animals, such as
pigs and humans. == Chemistry ==