The base of the Rupelian Stage (which is also the base of the Oligocene Series) is at the
extinction of the
foraminiferan genus Hantkenina. An official
GSSP for the base of the Rupelian has been assigned in 1992 (
Massignano, Italy). The transition with the Chattian has also been marked with a GSSP in August 2017 (
Monte Conero, Italy). The top of the Rupelian Stage (the base of the Chattian) is at the extinction of the foram genus
Chiloguembelina (which is also the base of foram
biozone P21b). The Rupelian overlaps the
Orellan,
Whitneyan and lower
Arikareean North American Land Mammal Ages, the upper
Mustersan and
Tinguirirican South American Land Mammal Ages, the uppermost
Headonian, Suevian and lower
Arvernian European Land Mammal Mega Zones (the Rupelian spans the
Mammal Paleogene zones 21 through 24 and part of 25), and the lower Hsandgolian
Asian Land Mammal Age. It is also coeval with the only regionally used upper Aldingan and lower Janjukian stages of Australia, the upper Refugian and lower Zemorrian stages of California and the lower Kiscellian
Paratethys stage of Central and eastern Europe. Other regionally used alternatives include the Stampian, Tongrian, Latdorfian and Vicksburgian. ==References==