The assignment of
Cigara to the Eocrinoidea has been based on the presence of respiratory structures known as epispires in middle section of the body.
Rejected affinities While
Cigara has previously been proposed as synonymous with
Dendrocystites (a
solute), or with
Acanthocystites (a
gogiid eocrinoid), the 1967
Treatise on Invertebrate Paleontology deemed these associations "unacceptable" and placed it in the section for unclassified eocrinoids. More recent research has suggested that the stylophoran aulacophore is most parsimoniously interpreted as a modified brachiole, as the water vascular system is not accompanied by the a somatocoelar derivative as it is in crinoid arms. Additionally, while most brachioles have biserial ossicles, in several species they are monoserial as they are in stylophorans. Therefore
Cigara having stylophoran affinites would not remove it from the blastozoa. Later taxonomies have continued to include
Cigara in Eocrinoidea, albeit tentatively. ==References==