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Columella (genus)

Columella is a genus of very small, air-breathing land snails, terrestrial pulmonate gastropod mollusks in the family Truncatellinidae, the whorl snails and allies.

Taxonomy
Columella is commonly classified within the Vertiginidae. However, as shown by Nekola & Coles (2016), the genus is not that closely related to the Vertiginidae, but may rather be affiliated with the Chondrinidae or form a pupilloid family of its own. A systematic revision is pending. ==Species==
Species
Species within the genus Columella include: • Columella acicularis Almuhambetova, 1979 • Columella aspera Waldén, 1966 • Columella columella (G. v. Martens, 1830) • Columella edentula (Draparnaud, 1805) - toothless column • Columella hartmutnordsiecki Schlickum & Geissert, 1980 • Columella hasta (Hanna, 1911) • Columella intermedia Schileyko & Almuhambetova, 1984 • Columella microspora (R.T. Lowe, 1852) • Columella nymphaepratensis Hlaváč & Pokryszko, 2009 • Columella polvonense (Pilsbry, 1894) • Columella simplex (Gould, 1840) • Columella talgarica Schileyko & Rymzhanov, 2010 • Columella wyciski Schlickum & Strauch, 1972 ;Species brought into synonymy: • Columella inornata (Michaud, 1831): synonym of Columella edentula (Draparnaud, 1805) (junior synonym) • Columella ninagongonis Pilsbry, 1935: synonym of Truncatellina ninagongonis (Pilsbry, 1935) (original combination) • Columella tridentata A. B. Leonard, 1946: synonym of Gastrocopta ruidosensis (Cockerell, 1899) ==References==
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