The Cedar Mountain Formation is proving to contain one of the world's richest and most diverse Early Cretaceous dinosaur faunas. The discoveries to date have revealed that the origin of some of the later Cretaceous dinosaurs may lie in Cedar Mountain, but further work is needed to understand the timing and effects the changing position of the North American Plate had on dinosaurian evolution. Also needed is a better understanding of the effects that the changing North American Plate had on the non-dinosaur vertebrates.
Dinosaurs The Cedar Mountain Formation is one of the last major dinosaur-bearing formations to be studied in the United States. Although sporadic bone fragments were known before 1990, serious research did not begin until that year. Since then, several organizations have conducted fieldwork collecting dinosaurs, chiefly the Oklahoma Museum of Natural History, the Denver Museum of Nature & Science, Utah State University-Eastern (formerly College of Eastern Utah), the Utah Geological Survey, Brigham Young University, and Dinosaur National Monument staff. This research indicates that at least two, possibly three dinosaur assemblages are contained within the formation. The oldest of these assemblages is from the Yellow Cat, Poison Strip and basal Ruby Ranch members. The small,
Ornitholestes-like
theropod Nedcolbertia and the brachiosaurid
sauropod Cedarosaurus may be considered as relics, with their closest relatives in the Morrison Formation. In contrast, the polacanthid
ankylosaur Gastonia and a yet unnamed
iguanodontid are similar to related forms from the Lower Cretaceous of southern England. These dinosaurs show that the connection between North America and Europe still existed during the Barremian. All of this changes, however, with the upper dinosaur assemblage from the top of the Ruby Ranch and Mussentuchit members. This upper assemblage shows greater similarities with Asian dinosaur assemblages from the same time. The upper assemblage also has a
tyrannosauroid, a
ceratopsian, and a
pachycephalosaur. Although not a dinosaur, the primitive mammal
Gobiconodon is known from both Mongolia and the Mussentuchit Member. Evidence for a middle dinosaur assemblage between the older and younger ones is controversial because the evidence mostly depends on a single specimen of the ornithopod
Tenontosaurus from high in the Ruby Ranch Member and the sauropod
Astrodon from low in the Ruby Ranch. Regardless, the upper and lower dinosaur assemblages in the Cedar Mountain Formation document the separation of North America and Europe, the westward drift of North America, and its connection with Asia 10 to 15 million years later. Data from Carpenter (2006), Cifelli
et al. (1999), Kirkland and Madsen (2007), and The Paleobiology Database.
Ankylosaurs Neornithischians A large sail-backed
iguanodont represented by large vertebrae and fragmentary remains from the Upper Yellow Cat Member.
Sauropods Theropods Indeterminate allosauroid material present in the Lower Yellow Cat, Upper Yellow Cat and Ruby Ranch Members. Indeterminate dromaeosaurine present in the Mussentuchit Member. Indeterminate deinonychosaurian remains present in the Mussentuchit member. Indeterminate velociraptorine remains present in the Mussentuchit Member. Indeterminate troodontid remains present in the Mussentuchit Member. Indeterminate therizinosaurid remains present in the Mussentuchit Member. Indeterminate dromaeosaurine remains present in the Mussentuchit Member. Possible indeterminate hesperornithiformes present in the Mussentuchit Member.
Other vertebrate fossils Besides dinosaurs, the Cedar Mountain Formation has produced a wealth of small fossils and microfossils, mostly teeth from a variety of vertebrates. Most of these specimens have been found in the Mussentuchit Member where they are collected by washing the rock through fine window screen. The teeth and other small fossils are picked from the residue. • Fish include primitive fresh or brackish water sharks (e.g.,
Hybodus) and rays (cf.,
Ischyrhiza), the lungfish (
Ceratodus) and several bony fishes known from vertebrae. Lungfish are able to breathe air when pond water becomes poorly oxygenated, such as during the dry season. • Amphibians include both salamanders (e.g.
Albanerpeton) and frogs, but neither is common. • Reptiles are more abundant and better studied. These include aquatic turtles (
Glyptops,
Naomichelys), at least one type of snake (
Coniophis), and several different lizards, including teiids (
Bicuspidon), possible
skinks and some extinct families (e.g., Paramacellodidae). Crocodiles are also present but their remains are fragmentary. They include
Bernissartia, an unnamed atoposaurid, and unnamed pholidosaurid. At least one fragment of a large pterosaur is known from the base of the Mussentuchut Member. Unfortunately, it is too incomplete to identify to family or genus. • Bird remains are very fragmentary because of their delicate structure. At least one aquatic bird is known. Based on the diversity of birds from the Early Cretaceous of China, other birds were probably present in Utah at this time as well. • Mammals are the most thoroughly studied thanks to the work of Jeffrey Eaton and Richard Cifelli. They include triconodonts (e.g.,
Astroconodon), which have the molar cusps arranged in a single row; symmetrodonts (e.g.,
Spalacolestes;
Spalacotheridium), characterized by molars having three cusps arranged in a triangle; multituberculates (e.g.,
Janumys;
Cedaromys;
Paracimexomys), with their multiple rows of cusps on the molars; one of the earliest marsupials (
Kokopellia), and several unnamed tribotheres, characterized by molars having three cusps that are typically asymmetrically arranged. The various vertebrates are listed by member in the list below. Non-vertebrate fossils are more widely distributed in the Cedar Mountain Formation. These include the distinctive reproductive structures of fresh water algae that are called
charophytes. Charophytes are so distinctive that they are used to correlate strata of similar age, and thus were used to show that the Yellow Cat Member was time equivalent to Barremian age strata in England.
Ostracods, small crustaceans with clam-like shells, also occur in fresh water deposits, along with "finger-clams" or conchostracans. Pollen have been found in the Mussentuchit Member and are important for reconstructing the environment. In a few places, large petrified logs are known, especially from the Poison Strip. These
conifer logs are over a meter in diameter and indicate the presence of trees over 30 m (100 feet). The distinct wood of the
tree fern Tempskya is occasional found as well. Data from Carpenter (2006), Cifelli
et al. (1999), Kirkland and Madsen (2007), and The Paleobiology Database.
Other reptiles Indeterminate isolated pterosaur remains have been recovered from the Yellow Cat and Mussentuchit Members. A partial
neochoristodere femur is known from the Yellow Cat Member.
Crurotarsans Indeterminate crocodilian remains present in the Yellow Cat and Ruby Ranch Members. Indeterminate pholidosaurid remains present in the Mussentuchit Member. Indeterminate atoposaurid remains present in the Mussentuchit Member.
Lepidosaurs Turtles Indeterminate baenid remains present in the Mussentuchit Member.
Amphibians Indeterminate anuran remains present in the Mussentuchit Member.
Fish Bony fish Indeterminate amiiform present in the Yellow Cat and Mussentuchit Members. Indeterminate neopterygian remains present in the Mussentuchit Member. Possible indeterminate pycnodontid remains present in the Mussentuchit Member. Possible indeterminate lepisosteid remains present in the Mussentuchit Member.
Cartilaginous fish A new genus and species of orectolobid present in the Mussentuchit Member.
Mammaliaformes New genus and species of pappotheriid present in the Mussentuchit Member. Indeterminate genus and species of picopsid present in the Mussentuchit Member. == See also ==