Physiography The Tuolumne River watershed can be divided into three distinct physiographic regions. From the headwaters to below Hetch Hetchy Reservoir, the river drains the Sierra highlands, which are characterized by solid,
glacier-carved granitic bedrock, thin poor soils and areas of heavy forestation. Most of the snowpack that feeds the Tuolumne accumulates in this region. Streams are generally high-gradient, clear and rocky, but large
alpine meadows provide riparian and wetland habitat, which contribute sediment to the watershed's runoff. There are also a large number of natural lakes and tarns in the upper watershed. The Sierra crest harbors a number of glaciers, including Lyell and Maclure Glaciers, the largest in Yosemite National Park. Melting ice feeds the upper reaches of the Tuolumne River, maintaining water flow in the late summer when many other streams in the region are dry. The Lyell Glacier is the second largest in the Sierra Nevada, after
Palisade Glacier at the headwaters of
Big Pine Creek a tributary of the
Owens River. Both Lyell and Maclure glaciers have shrunk significantly since the late 1800s when measurements were first made. Between Hetch Hetchy Reservoir and the upper end of Lake Don Pedro the Tuolumne flows through the Sierra foothills, where it has a lower gradient but continues to maintain a fast current. Stream pools and gravel bars figure more prominently in the channel, allowing the growth of riparian habitat. By the time the river reaches Lake Don Pedro it has accumulated almost 90 percent of its total flow as precipitation run-off,
snowmelt and
groundwater base flow. Below Don Pedro Dam the river flows across the large
alluvial plain of the San Joaquin Valley, formed by millions of years of sediment deposits eroded from the Sierra. Historically this section of the river was subject to frequent course changes and formed thousands of acres of seasonal wetlands during the spring floods. Since the introduction of large-scale agriculture the Tuolumne River course has been fixed between an extensive system of levees. More than in the San Joaquin Valley are irrigated using Tuolumne water. About , or 34 percent, of the Tuolumne River watershed is inside Yosemite National Park. The rivers whose watersheds border that of the Tuolumne are the
Stanislaus River and
West Walker River in the north, the
Merced River in the south, the
East Walker River to the northeast, the
Mono Lake basin (
Mill Creek,
Lee Vining Creek and
Rush Creek) to the east, and the headwaters of the San Joaquin River proper to the southeast.
Tributaries Major tributaries of the Tuolumne and pertinent data are listed in the below table. Order is from the headwaters downstream.
Land use Almost 80 percent of the Tuolumne River watershed lies above
New Don Pedro Dam and is primarily forested with some alpine, meadow and grassland zones. The primary land use in the lower Tuolumne River watershed is agriculture, but there are also significant urbanized areas. Although only about (5.4 percent) of the watershed itself are farmed, river water is used to irrigate almost five times that area outside the watershed boundaries. About (1.3 percent) of the watershed are urbanized, mostly in the Greater Modesto area. About 550,000 people inhabit the Tuolumne River watershed as a whole, with approximately 210,000 living in Modesto.
Engineering and flood control Like most other Sierra Nevada rivers, the Tuolumne has been extensively dammed to provide
flood control, water supply, and hydroelectric power. The uppermost dam is the O'Shaughnessy, which impounds the Hetch Hetchy Reservoir, providing water and power to the City of San Francisco. The Hetch Hetchy hydroelectric system diverts large volumes of water into tunnels that feed two hydroelectric plants, the Kirkwood and the Moccasin. The power system is efficient because of the Tuolumne's great drop, over , between Hetch Hetchy and the outlet of Moccasin Powerhouse. However, it also causes dewatering of significant portions of the Tuolumne River, especially during low water summers when all but a minimal fish release is diverted through the power plants. Not all of the flow used by the power plants is returned to the river. About per year flows through the gravity fed Hetch Hetchy Aqueduct to serve San Francisco and its customer municipalities. The previous time it was used was 1988. Below the confluence with the North Fork of the Tuolumne the river flows into Lake Don Pedro, the largest in the Tuolumne River system and the sixth-largest man made lake in California at . The earth-filled New Don Pedro Dam, high, is the 10th tallest dam in the United States and was constructed as a multipurpose project, serving irrigation, municipal water supply, hydropower and flood control. It replaced an earlier concrete dam built on approximately the same site. The Turlock and Modesto Irrigation Districts control the lion's share of water in the reservoir, although the City of San Francisco also has storage rights, which it uses mainly to offset the effects of water diverted upstream at Hetch Hetchy. Because of the desire to increase water storage for other uses, only of the reservoir is designated flood control, which is relatively small compared to other reservoirs in California. As a result, Don Pedro frequently fills and spills during winter storms, causing damage downstream. The La Grange Dam, a much smaller masonry diversion dam, is the last one on the Tuolumne before it joins with the San Joaquin. Modesto Canal, on the north side of the river, and Turlock Canal, on the south side, are both large irrigation diversions. The Modesto Canal feeds the off-stream Modesto Reservoir, which regulates the water for distribution to farmers. The Turlock Lake is regulated by the similar but much larger Turlock Lake, which has a capacity of . The estimated annual diversion for irrigation is , about half of the Tuolumne River's natural flow.
Fish, plants and wildlife The Tuolumne once supported large spring and fall runs of
chinook salmon (king salmon,
Oncorhynchus tshawytscha) and
steelhead trout (
Oncorhynchus mykiss). In its native state the fall Chinook run alone may have numbered as high as 130,000. With the exception of the
Merced River immediately south, this is the southernmost existing population of Chinook salmon in North America. Massive irrigation diversions and river channelization projects have led to the extinction of the spring Chinook run and greatly reduced the fall run. Dams have blocked more than half of the original of migratory fish spawning habitat in the Tuolumne River watershed. The San Joaquin River and the
Sacramento–San Joaquin River Delta, which link the Tuolumne salmon runs to the Pacific, have also been affected by flow reduction and pollution. In addition, introduced species such as
striped bass prey on juvenile salmon, consuming as much as 93 percent of smolt before they can migrate to the sea. A controversial proposal by the State Water Resources Control Board from around 2013 would require increased spring and summer flows on the Tuolumne River which would benefit native
anadromous fish populations, but greatly reduce the water supply available for farms and cities. This has been countered by irrigation districts who maintain that the aforementioned predation, not low flows, are the primary cause of salmon smolt deaths. Forests in the watershed are divided between several distinct zones. The foothill region is characterized by trees and plants adapted to a hot, dry climate, including
chamise (greasewood),
California lilac,
manzanita,
blue oak,
interior live oak (Sierra live oak), and
gray pine. Further east and higher in elevation are the lower and upper montane forests, which lie between and elevation, respectively. The lower montane forests include
black oak,
ponderosa pine,
incense cedar and
white fir; higher up
Jeffrey pine and
western juniper appear. Higher still is the subalpine forest zone between , which is dominated by
Western white pine,
mountain hemlock and
lodgepole pine. Above 9500 feet, only small shrubs, flowering plants and occasional
whitebark pine populate the alpine zone, which is limited to the peaks around Tuolumne Meadows and along the Sierra Crest. Humans have influenced natural conditions in the Tuolumne River watershed for centuries. Native Americans set
controlled fires in grassland to clear out dead foliage, providing room for new growth which in turn attracted game animals to forage. They also periodically burned out oak woodland areas in the foothills to control pests, which otherwise would have affected their main food supply, acorns. During the Spanish and Mexican periods, the Central Valley was used for cattle ranching, which quickly decimated natural range grasses. During the 1880s American sheep ranchers brought their herds into the upper Tuolumne River watershed for summer pasture, which further damaged the range in the foothills and Sierras. Hetch Hetchy Valley was one of the main summer grazing areas for sheep. The decline of native
perennial grasses caused invasive
annual species to flourish, and also increased the susceptibility of the watershed to
wildfires. In order to protect property and watershed quality, intensive fire suppression began in the early 20th century. However, this led to overgrowth and a disruption of the natural
forest succession process that was previously driven by wildfires. ==Early history==