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Rachel Bloom as Rebecca Bunch, a lawyer, originally from
Scarsdale, New York. Rebecca suffers symptoms of depression and
anxiety and is prone to impulsive decisions and delusions, and she tends to
dissociate when she is under emotional distress. Rebecca uproots her life and relocates to the suburb of West Covina, California, in an attempt to win back her crush, Josh Chan, whom she briefly dated in a summer camp ten years prior. Intelligent but often lacking awareness of appropriate behavior, Rebecca comes across as unpredictable, selfish, self-centered, needy, and eccentric to those around her. At the same time, she often tries hard to be kind and thoughtful, and she makes loyal friends in West Covina. In the third season, she is diagnosed with
borderline personality disorder (BPD) and makes positive steps towards managing her mental health and, in the fourth season, she quits her job as a lawyer, realizing it makes her unhappy, and becomes involved in songwriting. •
Vincent Rodriguez III as Josh Chan, the object of Rebecca's affections for much of the series. Josh demonstrates commitment issues in his relationship with Rebecca and is an oblivious person. Like Rebecca, Josh has a tendency to be self-centered and immature. Josh later improves as a person and apologizes to Rebecca for the mistakes he made in their relationship. He subsequently goes on a journey of self discovery as he tries to find his path and understand what he wants in life. •
Santino Fontana (seasons 1–2) and
Skylar Astin (season 4) as Greg Serrano, a smart but underachieving bartender and Josh's best friend, who has complicated feelings for Rebecca. He departs West Covina to attend business school at
Emory University, partly as a way to get away from his destructive relationship with Rebecca, despite his feelings for her. Although he struggles with alcoholism throughout season 1, he comes to terms with his addiction in season 2. He returns to West Covina in season 4, more emotionally healthy and played by a different actor. •
Donna Lynne Champlin as Paula Proctor (née O'Brien), Rebecca's co-worker and new best friend. As a way to distract herself from her own failing marriage, Paula hatches schemes and gives questionable advice in support of Rebecca's pursuit of Josh, mostly to compensate for her lack of fulfillment from not pursuing her dreams when she was younger. She later begins to focus instead on pursuing her law degree and establishes healthier boundaries with Rebecca, distancing herself from her past scheming. She and her husband Scott tend to be neglectful parents to their two sons. •
Pete Gardner as Darryl Whitefeather, Rebecca's sentimental, emotional and often clueless boss. A middle-aged divorced dad, he discovers he is
bisexual and begins dating White Josh. He is prone to feelings of loneliness and neediness over the fact that he originally lacked strong friendships and real personal connections with others. He later forms friendships with his colleagues Maya, Nathaniel, Rebecca, and Paula, whom he considers his best friend. He and White Josh break up in the third season due to Darryl's desire to have a baby, but remain good friends. He has a baby girl carried by Heather with Rebecca's egg and names her Hebecca. Although he has a good heart, Darryl's neediness and persistence can push people away from him. In Season 4, he marries a woman named April who's the mother of his daughter's enemy. •
Vella Lovell as Heather Davis, Rebecca's cool college student neighbor and sometimes roommate. She is apathetic and sardonic, and she lacks motivation for obtaining achievement or self-sufficiency, because her parents encouraged failure and coddled her. In season 3, Heather is forced by her college to graduate, despite her wishes; she becomes a regional manager for three Home Base bars and Darryl's surrogate mother. She briefly dates Greg in season 1; in season 3, she starts a relationship with Hector, whom she eventually marries. •
Gabrielle Ruiz as Valencia Perez (seasons 2–4; recurring season 1), Josh's controlling girlfriend and Rebecca's rival for Josh's affections. She initially had a strong resistance to forming relationships with women, as she saw them as competition, making her disliked by her female peers. She has insecurity about her body image and can be condescending to people who she feels don't meet her standards. After breaking up with Josh, she questions her life choices and feels she lacks fulfillment, having devoted 15 years of her life to Josh. After Rebecca also breaks up with Josh, she and Valencia become close friends. Valencia later meets and falls in love with a woman named Beth, with whom she runs a party planning business. They get engaged by the end of Season 4. •
David Hull as Josh Wilson (season 3; recurring seasons 1–2; special guest season 4), Josh Chan's friend, a laid-back fitness instructor nicknamed "White Josh" to differentiate him from Josh Chan, who is of Filipino descent. He begins dating Darryl after the latter comes out as bisexual. White Josh tends to be judgmental, yet he is one of the most level-headed, rational characters in his social sphere. He and Darryl break up over Darryl's desire to have a baby, but stay on friendly terms. •
Scott Michael Foster as Nathaniel Plimpton III (seasons 3–4; recurring season 2), a wealthy and successful lawyer and Rebecca's new boss in season 2 after he buys equity in Whitefeather & Associates. He and Rebecca have a complicated relationship due to mutual dislike yet intense physical attraction to one other, and they explore a relationship through part of season 3. He possesses an
inferiority complex, feeling that he needs to live up to his father's expectations; he tends to become self-loathing and defeatist when he fails to do so. Nathaniel exhibits some
sociopathic tendencies; for example, he has suggested the murder of another character's relative as revenge. After experiencing feelings of loss and abandonment after his breakup with Rebecca, he resolves to become a kinder person. Similarly to Rebecca, he quits being a lawyer for MountainTop and starts legally representing a zoo in Season 4. ==Episodes==