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Cretalamna

Cretalamna is a genus of extinct otodontid shark that lived from the latest Early Cretaceous to Eocene epoch. It is considered by many to be the ancestor of the largest sharks to have ever lived, such as Otodus angustidens, Otodus chubutensis, and Otodus megalodon.

Taxonomy
Research history s of Otodus appendiculatus (Fig. 1-25) and holotype of C. lata (Fig. 26) in the third volume of Recherches sur les poissons fossiles; the tooth in Fig. 10 is the sole lectotype of C. appendiculata|left|upright=1.2 Cretalamna was first described by Swiss naturalist Louis Agassiz using five teeth previously identified as the common smooth-hound and collected by English paleontologist Gideon Mantell from the Southerham Grey Pit near Lewes, East Sussex. In his 1835 publication Rapport sur les poissons fossiles découverts en Angleterre, he reidentified them as a new species of porbeagle shark under the taxon Lamna appendiculata. The species would later be found in 1958 by Soviet paleontologist Leonid Glickman to belong to a distinct new genus- Cretalamna. Despite Agassiz's remarks on variability, promoted into a subspecies in 1977 by Belgian paleontologist Jaques Herman, and finally elevated to the species level as Cretolamna lata by Herman and paleontologist Van Waes Hilde in 2012. In 1897, French paleontologist Fernand Priem described a single tooth from the Köpinge Sandstone in Scania, Sweden under the taxon Lamna borealis. This would be revised to '' 'Cretolamna borealis' by Glickman in a 1980 paper. In 1935, French Paleontologist Camille Arambourg described a new subspecies of C. biauriculata from teeth found in Moroccan phosphates under the taxon Lamna biauriculata maroccana, which was elevated into its own species in 1997. In 1972, French ichthyologist Henri Cappetta described teeth from Maastrichtian deposits near the Mentès well in Tahoua, Niger, which he assigned to the subspecies Lamna biauriculata nigeriana. This subspecies would also be elevated to its own species in 1991. In 1975, Cappetta and American paleontologist Gerard Case examined Cretalamna teeth described by Arambourg in 1952 from Danian deposits in Morocco and proposed that it represents a new subspecies of the type species and assigned it the taxon Cretolamna appendiculata arambourgi, which Siversson et al.'' (2015) elevated into its own species. Etymology The genus Cretalamna is a portmanteau of creta, the Latin word for "chalk", prefixed to the genus Lamna, which is a romanization of the Ancient Greek λάμνα (lámna, meaning "kind of fierce shark"). When put together they mean "chalk-shark", which refers to chalk deposits from which the species' type specimens were found in. C. biauriculata's specific epithet is a portmanteau derived from the Latin prefix bi- (two) prefixed onto the Latin auriculātā (eared), together meaning "having two ears". This is a reference to the species' large lateral cusplets, which somewhat resemble a pair of ears. Since then, the reinstatement of '' 'Cretalamna' gained prominence and by the 2010s, was accepted by the majority of paleontologists. However, some paleontologists including Cappetta strongly opposed it. In an attempt to suppress the usage of 'Cretalamna' , Cappetta appealed to a representative of the ICZN, arguing that the original intentions of Glickman and the prevailing usage of 'Cretolamna' '' prior to Siversson (1999) secures its priority. The ICZN, who reportedly were impressed by Cappetta's "spirit", subsequently erected Article 33.3.1 of the 2000 Edition of the Code in order to address this situation in the future, which states that "when an unjustified emendation is in prevailing usage and is attributed to the original author and date it is deemed to be a justified emendation". While Cappetta argued in a 2012 handbook that this new provision justifies the priority of '' 'Cretolamna' '' due to the spelling's overwhelmingly prevailing usage prior to its replacement by Siversson in 1999, Siversson himself pointed out in a 2015 paper that the provision cannot be worked retroactively, and that the continued prevailing usage of '' 'Cretalamna' '' since the provision's establishment ironically secures its priority rather than threaten it. '' 'Cretalamna' currently remains as the most prevalent spelling and paleontologists have expressed the unlikeliness of a return to the usage of 'Cretolamna' ''. ==Description==
Description
or salmon shark-like profile Cretalamna was a medium to large-sized shark. Based on vertebral comparisons with various extant lamniforms and Cretoxyrhina, a 2007 study by Kenshu Shimada estimated a total length of for the most complete skeleton of a large individual (LACM 128126; C. hattini holotype Subsequently, in 2019, the teeth of C. appendiculata from Himedo Park, Kugushima and Wadanohana which are larger than those in LACM 128126 yielded maximum length estimates of up to , and , respectively. It was most similar to the porbeagle and salmon sharks in build, with a compact fusiform body, large pectoral and first dorsal fins and tail, and small second dorsal, pelvic, and anal fins. The first dorsal fin was positioned directly above the pectoral fins unlike its analog species, where the first dorsal fin is usually positioned behind it. Such a body plan is indicative of an active fast-swimming pelagic shark likely partially warm-blooded through regional endothermy. In their 2024 study's appendix, Sternes and colleagues questioned this particular specimen's authenticity and anatomical parts which can only be speculated by photographs and its uncertain catalog status did not allow the reproducibility of the proportions suggested by Greenfield (2022). Dentition (Morocco) Cretalamna teeth are distinguished by a broad triangular cusp and two lateral cusplets. The cutting edges of the teeth are razor-like, while the sides have a smooth surface. Teeth symmetry is variable; some have exact bilateral symmetry whereas others have high asymmetry. Adjacent teeth do not overlap. Jaw In C. hattini, the upper and lower jaws are similar to that of Cretoxyrhina mantelli. The jaws also resemble those of modern alopiids (thresher sharks) and lamnids. Limited fossil evidence suggests that the upper jaws extended over the lower jaws, giving Cretalamna a subterminal mouth. == Paleoecology ==
Paleoecology
Distribution Cretalamna was a widespread genus found in North Africa (Morocco), the Near East (Jordan), West Africa (Mali), North America both on the East Coast and in the Midwest and Central America (Tonosí, Panama). Deposits in Morocco are usually Eocene in age; deposits in Jordan are of Cretaceous and Eocene in age; most deposits in the U.S. are of Cretaceous and Paleocene age; and deposits in Mali are of Cretaceous (Maastrichtian) age. The Cretaceous waters inhabited by Cretalamna were also home to a diverse range of cartilaginous fishes, bony fishes, turtles, squamates, plesiosaurs, pterosaurs, birds, and even some non-avian dinosaurs. For example, multiple teeth of C. appendiculata have been found around elasmosaurid Futabasaurus, suggesting it predated or scavenged that elasmosaur. Some tooth specimens of Cretalamna exhibit heavy wear—likely the result of drastic diet changes. Extinction A possible factor to the extinction of Cretalamna is increased competition with newer generalist sharks during the Cenozoic. It is widely believed that Otodus is derived from Cretalamna due to its strong similarity to certain species within the genus. == See also ==
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