He received his
A.B. in
botany, and an
M.A. and
Ph.D. in
paleobotany from the
University of California at Berkeley. He served in the
United States Army during
World War II performing strategic analysis of
aerial photographs of
terrane. After the war he was hired as an assistant professor of
geology at the
University of California at Los Angeles. He eventually became a full professor of both geology and
botany at UCLA before moving to the
University of California at Davis as a professor of paleoecology late in his career. He became professor
emeritus at Davis in 1976. He was elected a Fellow of the
American Academy of Arts and Sciences in 1981. His collections of fossil type floras are housed at the
University of California Museum of Paleontology == Awards ==