Lloyd's major contribution to
botany was in the field of
plant reproduction. His contributions to the field include a mechanistic treatment of different modes of
self-pollination in
hermaphroditic plants, a genetically defined continuum of plant gender, early development of theory of the evolution of separate sexes in plants, and with C.J. Webb, a challenge to conventional views of the evolution of
heterostyly. Because of his ideas and work on population biology of plants, he is sometimes referred to as the "
W.D. Hamilton in plant biology". ==References==