The fungus
over-winters as
pycnidia on the outside of diseased wood. The pycnidia produce and release dark brown, striated
conidia. The conidia are then dispersed by wind or rain-splash, spreading the fungi from one part of the vine to another, and to other nearby vines. Disease develops when conidia land on freshly cut or damaged wood; the conidia germinate in the wood tissue, progressively damaging the host-plant's vascular system. As tissue damage advances around the initial site of infection, the formation of
cankers and cell death - or
necrosis - will cause eventual dieback of the wood. Under certain conditions,
pseudothecia will grow on the surface of cankers and produce
ascospores, which - like conidia - will disperse to infect surrounding wounds. ==Management==