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Dorudon

Dorudon ("spear-tooth") is a genus of extinct basilosaurid ancient whales that lived alongside Basilosaurus 41.03 to 33.9 million years ago in the Eocene. It was a small whale, with D. atrox measuring 5 metres (16 ft) long and weighing 1–2.2 metric tons. Dorudon lived in warm seas around the world and fed on small fish and mollusks. Fossils have been found along the former shorelines of the Tethys Sea in present-day Egypt and Pakistan, as well as in the United States, New Zealand and Western Sahara.

Taxonomic history
described Dorudon serratus based on a fragmentary maxilla and a few teeth found in South Carolina. He concluded that the teeth must have belonged to a mammal since they were two-rooted, that they must have been teeth from a juvenile since they were hollow, and also noted their similarity to the teeth then described for Zeuglodon (Basilosaurus). When exploring the type locality, Gibbes discovered a lower jaw and twelve caudal vertebrae, which he felt obliged to assign to Zeuglodon together with his original material. Gibbes concluded that Dorudon were juvenile Zeuglodon and consequently withdrew his new genus. He did, however, allow Louis Agassiz at Harvard to examine his specimens, and the Swiss professor replied that these were neither teeth of a juvenile nor those of Zeuglodon, but of a separate genus just as Gibbes had first proposed. described Prozeuglodon atrox (="Proto-Basilosaurus") based on a nearly complete skull, a dentary and three associated vertebrae presented to him by the Geological Museum of Cairo. however, realized that Andrews' specimen was a juvenile, and, he assumed, the same species as Zeuglodon isis, described by Andrews 1906. Kellogg also realized that the generic name Zeuglodon was invalid and therefore recombined it Prozeuglodon isis. Since then many specimens have been referred to Prozeuglodon atrox, including virtually every part of the skeleton, and it has become obvious that it is a separate genus, not a juvenile "Proto-Zeuglodon". Kellogg placed several of the species of Zeuglodon described from Egypt in the early 20th century (including Z. osiris, Z. zitteli, Z. elliotsmithii and Z. sensitivius) in the genus Dorudon. synonymized these four species and grouped them as Saghacetus osiris. D. serratus from Georgia and South Carolina in the United States. The type species D. serratus was, and still is, based solely on two partial maxillae with a few teeth, cranial fragments, and a dozen vertebrae with some additional material, collected but not described by Gibbes, and referred to the type species. Before Uhen 2004, D. atrox was based solely on Andrews holotype skull, lower jaw, and the vertebrae he referred to it, but is now the best-known archaeocete species. --> The two species of Dorudon differ from other members of Dorudontinae mainly in size: they are considerably larger than Saghacetus and slightly larger than Zygorhiza, but also differ from both these genera in dental and/or cranial morphology. The limited known material for D. serratus makes it difficult to compare the two species of Dorudon. placed D. atrox in the same genus as D. serratus because of similarities in size and morphology, but kept them as separate species because of differences in dental morphology. Even though D. serratus is the type species, the description of Dorudon is largely based on D. atrox because of its completeness. The cranial morphology of D. atrox makes it distinct from all other archaeocetes. ==Description==
Description
, Washington, D.C. Dorudon was a medium-sized whale, with D. atrox reaching in length and in body mass. Dorudontines were originally believed to be juvenile individuals of Basilosaurus as their fossils are similar but smaller. They have since been shown to be a different genus with the discovery of Dorudon juveniles. Although they look very much like modern whales, basilosaurines and dorudontines lacked the melon organ that would allow their descendants to use echolocation as effectively as modern whales. Like other basilosaurids, their nostrils were midway from the snout to the top of the head. Dentition The dental formula for Dorudon atrox is . Typical for cetaceans, the upper incisors are aligned with the cheek teeth, and, except the small I1, separated by large diastemata containing pits into which the lower incisors fit. The upper incisors are simple conical teeth with a single root, lacking accessory denticles, and difficult to distinguish from lower incisors. The upper incisors are missing in most specimens and are only known from two specimens. The upper canine is a little larger than the upper incisors, and, like them, directed slightly buccally and mesially. The lower premolars are double-rooted, buccolingually compressed teeth, except the deciduous P1 which is single-rooted. P3 is the second-largest cheek tooth, P4 the largest; both are very similar, dominated by the central cusp. == Paleoecology ==
Paleoecology
Dorudon calves may have fallen prey to hungry Basilosaurus, as shown by unhealed bite marks on the skulls of some juvenile Dorudon. ==See also==
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