Douglas Koshland

Koshland is the son of Marian (née Elliot) and Daniel E. Koshland Jr. He earned his B.A. in chemistry from Haverford College and his Ph.D. in microbiology from Massachusetts Institute of Technology under the guidance of David Botstein. He then did his postdoctoral work with Leland Hartwell at University of Washington and with Marc Kirschner at the University of California, San Francisco. He was a staff scientist at Carnegie Institution for Science from 1987 and an adjunct professor in the Department of Biology at Johns Hopkins University. He was a Howard Hughes Medical Institute (HHMI) investigator from 1997-2012, was inducted into the National Academy of Sciences in 2010, and is a Fellow of the American Association for the Advancement of Science. ==Personal life==

Koshland is married to Mary Porter. == References==