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Eocene Okanagan Highlands

The Eocene Okanagan Highlands or Eocene Okanogan Highlands are a series of Early Eocene geological formations which span a 1,000 km (620 mi) transect of British Columbia, Canada, and Washington state, United States. Known for a highly diverse and detailed plant and animal paleobiota the paleolake beds as a whole are considered one of the great Canadian Lagerstätten. The paleobiota represented are of an upland subtropical to temperate ecosystem series immediately after the Paleocene–Eocene thermal maximum, and before the increased cooling of the middle and late Eocene to Oligocene. The fossiliferous deposits of the region were noted as early as 1873, with small amounts of systematic work happening in the 1870–1920s on British Columbian sites, and 1920–1930s for Washington sites. Focus and more detailed descriptive work on the Okanagan Highland sites started in the late 1960s.

Extent
The majority of the paleolake deposits are compression fossils in lake bed sediments spanning a transect, The Northern sites are composed of unnamed Ootsa Group formations which outcrop as the "Driftwood shales" near Smithers, British Columbia, and the "Horsefly shales", of an unnamed formation and unnamed group which outcrop around Horsefly, British Columbia. There is debate as to the affiliation of the, now lost, Quesnel sites with the Greater Okanagan Highlands. Archibald et al (2018) in the monograph overview of the Highlands Hymenoptera included them as part of the series. However the certainty for the placement was later questioned by Archibald and Cannings (2022) who opted to tentatively exclude Quesnel from the highlands while discussing the history of field collecting in the region. ==History==
History
in 1885 The earliest geological work in south and central British Columbian sites was during exploratory expeditions under the leadership of George Mercer Dawson in the mid-1870s to document the coal and mineral resources of British Columbia. During the 1875 field work, fossil collections were made in the Quesnel region. Two years later, in 1877, the expedition explored the Okanagan, Nicola, and North Thompson valleys with field collecting along areas of the Similkameen River, Whipsaw Creek and Nine-Mile Creek, north at around Quilchena, and at several areas near Kamloops. These expeditions resulted in a series of papers on the plants, authored by John William Dawson, later David Pearce Penhallow and Edward Wilber Berry. The insects were first detailed by Samuel Hubbard Scudder, with follow-up papers by Anton Handlirsch. While reporting on additional plant fossils collected from British Columbia, Penhallow (1906) noted the likely coeval status of the Princeton basins with many of the sites now considered the Okanagan Highlands. The first brief work on fish from the Highlands occurred with the 1893 report by Edward Drinker Cope on several fish sent to him by George Dawson from the Tranquille and Princeton areas. While Cope deemed the Tranquille specimens too incomplete to identify, he did provide the description for "Amyzon" brevipinne from the Similkameen River fossils. in 1998 Republic, Washington, area fossils were first reported by Joseph Umpleby (1910), based on fish he collected near the Tom Thumb mine, and given a tentative late Miocene age. Two of the fish were figured and briefly mentioned 7 years later by Charles R. Eastman, who assigned them tentatively to "Amyzon" brevipinne, making one of the early connections between Republic and the British Columbian sites. Report of the plant fossils of the Republic area were first published by Berry (1929) who included the Republic fossils as part of the Latah Formation. The inclusion within an expanded Latah Formation was questioned by Roland W. Brown in 1936, who noted the similarities between Republic and other older fossil sites, combined with the Republic lake bed's overlying basalts then thought to be of similar age to the Columbia River Basalts. This similarity was again noted by Chester A. Arnold during his review of the conifer flora associated with the Princeton basin. Arnold noted the Allenby sites shared over half of the taxa that had been previously reported from Republic. Starting in the early 1960s and extending through the 1980s were a series of papers on the British Columbian sites combining palynology and the newly devised process of potassium–argon dating to better understand the geochronology of the sites. The term "Okanagan Highlands" for Eocene formations of the region was coined by Wesley Wehr and Howard Schorn in a 1992 Washington Geology paper on the conifer research at Republic. The name was derived from the current Okanagan Highlands but applied to the, as then identified, microthermal forests preserved at Republic and Princeton. The term was expanded upon, and by 2005 it was generally understood to encompass all Eocene fossiliferous formations between Republic and Smithers, British Columbia. The highlands as a whole have been described as one of the "Great Canadian Lagerstätten" ==Preservation types==
Preservation types
The majority of formations in the Eocene Okanagan Highlands preserve compression fossils in sandstone to shale rock deposited from lake environments where seasonal mixing and anoxia were prevalent. Additionally, two important non-compression biotas are present in the Eocene Okanagan Highlands: A permineralized chert flora, the Princeton Chert is found along the Similkameen River interbedded with coal deposits of the Ashnola shale unit, Allenby Formation, known for anatomically preserved plants. In the Central sites, subbituminous coal of the Hat Creek Coal Formation around Hat Creek hosts an amber fauna, the Hat Creek amber which preserves many small insects that would likely not be found in the compression biotas. ==Paleoclimate==
Paleoclimate
over Lake Edward Based on paleobotanical proxies and geochemical data, the highlands are thought to have been between in elevation, placing them similar to or higher than the modern sites. Estimates of the mean annual temperature have been derived from climate leaf analysis multivariate program (CLAMP) analysis and leaf margin analysis (LMA) of both the Princeton and Republic paleofloras. The CLAMP results, after multiple linear regressions for Republic, gave a mean annual temperature of approximately , while the LMA gave . Princeton's multiple linear regression CLAMP results gave a slightly lower , and the LMA returned a mean annual temperature of . This is lower than the mean annual temperature estimates given for the coastal Puget Group, which is estimated to have been between . The bioclimatic analysis for Republic and Princeton suggest mean annual precipitation amounts of and respectively. The warm temperate uplands floras of the highlands, associated with downfaulted lacustrine basins and active volcanism are noted to have no exact modern equivalents, due to the more seasonally equitable conditions of the Early Eocene. However, the highlands have been compared to the upland ecological islands in the Virunga Mountains within the Albertine Rift of the African rift valley. ==Paleobiota==
Paleobiota
The Eocene Okanagan Highlands hosted a diverse mix of temperate and tropical paleobiotic elements, The paleobotanical community was a mixture of plants found in subtropical evergreen and temperate deciduous forests. ==References==
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