About one-third of all fern species are epiphytes. The third largest group is
clubmosses, with 190 species, followed by a handful of species in each of the
spikemosses, other ferns,
Gnetales, and
cycads. Within the
Neotropics, there are estimated to be total of 15,500 species of epiphytic vascular plants. A monograph on epiphytic
plant ecology was written by
A. F. W. Schimper (, 1888). Assemblages of large epiphytes occur most abundantly in moist
tropical forests, but mosses and lichens occur as epiphytes in almost all biomes. In Europe there are no dedicated epiphytic plants using roots, but rich assemblages of mosses and lichens grow on trees in damp areas (mainly the western coastal fringe), and the common
polypody fern grows epiphytically along branches. Rarely, grass, small bushes or small trees may grow in
suspended soils up trees (typically in a rot-hole).
Holo-epiphyte or hemi-epiphyte Epiphytes can generally be categorized into holo-epiphytes or hemi-epiphytes. A holo-epiphyte is a plant that spends its whole life cycle without contact with the ground and a hemi-epiphyte is a plant that spends only half of its life without the ground before the roots can reach or make contact with the ground.
Orchids are a common example of holo-epiphytes and
strangler figs are an example of hemi-epiphytes.
Plant nutrient relations Epiphytes are not connected to the soil, and consequently must get nutrients from other sources, such as fog, dew, rain and mist, or from nutrients being released from the ground rooted plants by decomposition or leaching, and dinitrogen fixation. Epiphytes can have a significant effect on the microenvironment of their host, and of ecosystems where they are abundant, as they hold water in the canopy and decrease water input to the soil. Some non-vascular epiphytes such as lichens and mosses are well known for their ability to take up water rapidly. Epiphytes create a significantly cooler and more moist environment in the host plant canopy, potentially greatly reducing water loss by the host through transpiration.
Plant metabolism CAM metabolism, a water-preserving metabolism present among various
plant taxa, is particularly relevant to epiphytic communities. For example, it is estimated that among epiphytic orchids, as many as 50% are likely to use it. Other relevant epiphytic families which display such metabolism are
Bromeliacee (e.g. in genera
Aechmea and
Tillandsia),
Cactaceae (e.g. in
Rhipsalis and
Epiphyllum) and
Apocynaceae (e.g. in
Hoya and
Dischidia). ==Marine epiphytes==