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Glossary of spider terms

This glossary describes the terms used in formal descriptions of spiders; where applicable these terms are used in describing other arachnids.

Terms
AAbdomen or opisthosoma: One of the two main body parts (tagmata), located towards the posterior end; see also Abdomen § Other animalsAccessory claw: Modified at the tip of the in web-building spiders; used with to grip strands of the web • Anal tubercle: A small protuberance (tubercule) above the through which the anus opens • Apodeme: see • Apophysis (plural apophyses): An outgrowth or process changing the general shape of a body part, particularly the appendages; often used in describing the male : see • Atrium (plural atria): An internal chamber at the entrance to the in female haplogyne spiders BBidentate: Having two • Book lungs: Respiratory organs on the ventral side (underside) of the , in front of the , opening through narrow slits; see also Book lungs • Branchial operculum: see • Bulbus: see CCalamistrum (plural calamistra): Modified setae (bristles) on the of the fourth leg of spiders with a , arranged in one or more rows or in an oval shape, used to comb silk produced by the cribellum; see also CalamistrumCaput (plural capita): see • Carapace: A hardened plate (sclerite) covering the upper (dorsal) portion of the ; see also CarapaceCarpoblem: The principal on the male ; also just called the tibial apophysis • Cephalic region or caput: The front part of the , separated from the thoracic region by the • Cephalothorax or prosoma: One of the two main body parts (tagmata), located towards the anterior end, composed of the head ( or caput) and the thorax (thoracic region), the two regions being separated by the ; covered by the and bearing the , legs, and mouthparts • Cervical groove: A shallow U-shaped groove, separating the and thoracic regions of the • Chelate: A description of a where the closes against a tooth-like process • Chelicera (plural chelicerae): One of two appendages at the front of the , made up of basal portion, the , and the ; sometimes called the jaw; see also CheliceraeCheliceral furrow: A shallow groove on the basal portion of a accommodating the , usually having on its margins • Cheliceral tooth: A tooth-like extension on the margin of the • Chilum: A small hardened plate (sclerite) at the base of the , under the • Claw: see • Claw tuft: A dense group of hairs or bristles (setae) underneath the paired , usually well developed in hunting spiders is covered with red scales. • Clypeus: The area of the between the anterior (frontmost) and the anterior edge of the carapace • Colulus: A short protuberance in the middle of the underside of the in front of the , considered to be a modification of the • Conductor: see • Copulatory duct: An internal tube (duct) from the through which sperm enters the female; separate from the duct through which fertilized eggs pass in entelegyne spiders • Copulatory opening: An opening in the ventral of female spiders; in entelegyne spiders, a double opening in the through which the is inserted; in haplogyne spiders, a single opening through which male is inserted • Coxa: see • Crenulate: Having longitudinal ridges • Cribellum: A sieve-like plate in front of the , used in conjunction with the ; spiders with a cribellum are called cribellate, those without ecribellate; see also CribellumCuspule: A small spiny outgrowth ("wart") on the and of MygalomorphaeCymbium (plural cymbia): The end part of the of the in a mature male, usually hollowed out and bearing the DDionychous: With two on the of each leg; a feature of spiders in the clade DionychaDorsal groove: see • Dorsum: The upper (dorsal) portion or surface of the body or ; the adjective dorsal may be applied to the upper portion or surface of any part of the body; see EEcribellate: see • Embolus: see • Endite: see • Endosternite: An internal hardened plate (sclerite) • Entelegyne: A spider whose female has an and separate ducts leading to for sperm storage and to the uterus for fertilization, creating a "flow-through" system; see ; see also Entelegynae , viewed from above • Epigastric furrow or epigastric fold: A transverse slit towards the front (anterior) of underside of the ; the front pair of open at the edge of this furrow as do the genital openings () • Epigyne or epigynum (plural epigynes): A hardened plate on the underside of the female in which the are located; only fully developed in mature females of spiders; see also EpigyneEyes: The basic number of eyes is eight, typically arranged in two rows (e.g. as in Gnaphosidae); the front row are the anterior eyes, the row behind the posterior eyes; the four eyes to the edges are the lateral eyes, the four eyes in the centre the median eyes; the anterior median eyes are called the main eyes or direct eyes, while the other eyes are called the secondary eyes or indirect eyes; the number of eyes, their sizes and arrangement varies widely and is characteristic of spider families; see , FFalx (plural falces): A dated term for • Fang: The final hinged part of the , normally folded down into a groove in the basal part of the chelicera; venom is injected via an opening near the tip of the fang • Femur: see • Fertilization duct: A duct in female spiders leading from the to the uterus • Folium: A broad leaf-like marking along the medial line of the top of the • Fossa (plural fossae): A pit or depression, typically in the • Fovea (also called thoracic furrow or dorsal groove): A depression or pit in the centre of the of a spider marking an inward projection of the exoskeleton to which stomach muscles are attached GGenital opening: see • Gnathocoxa: see • Gonopore: The genital opening; located in the epigastric furrow; the opening of the duct from the uterus in females and from the testes in males; see also GonoporeGravid: A fertilised female with an enlarged abdomen section right before egg laying has begun. HHaematodocha (also spelled hematodocha): see • Haplogyne: A spider whose female lacks an and in which the same ducts are used to transport sperm to the uterus and to the ; see ; see also HaplogynaeHeart mark: A narrow marking along the top of the roughly corresponding to the location of the heart LLabio-sternum mound: A mound separating the from the , found in some tarantulas, where it can be a diagnostic feature • Labium (plural labia): A hardened plate (sclerite) between the at the front of the ; see also Arthropod mouthparts: LabiumLabrum: see • Lamella characteristica or just lamella: a sheet-like on the of many Linyphiidae species • Lateral (applied to appendages): Viewed from above or below, the sides of the leg or , i.e. the surfaces parallel to the line of sight; see , • Laterigrade: With legs directed to the side, hence appearing like and moving like a crab; see • Leg formula: The legs are numbered from the front from I to IV; the relative length of the legs can be represented by four numbers from the longest to the shortest; e.g. 1423 = first leg (leg I) is longest and third leg (leg III) is shortest • Leg parts or segments: see • Lorum: A set of covering the dorsal surface of the . MMain eye: One of the two anterior median eyes (AME) that have the light-detecting units (rhabdomeres) pointing towards the source; particularly enlarged in the families Salticidae and Thomisidae; see , • Mastidion (plural mastidia): A projection or bump on the chelicerae (not to be confused with ) • Maxilla (plural maxillae; also called endite or gnathocoxa): Modified of the , used in feeding; not the structure called by this name in other arthropods, for which see Maxilla (arthropod mouthpart)Metatarsus: see OOcular area or ocular quadrangle: The area of the which includes the . • Operculum or branchial operculum (plural opercula): One of the plates on the surface of the , just in front of the , covering the , often pale, yellow or orange in colour; two pairs in Mygalomorphae, one pair in other spiders • Opisthosoma: see PPalp: see • Palpal bulb (also called bulbus, palpal organ, genital bulb): The copulatory organ of the male spider, carried on the modified last segment of the , used to transfer sperm to the female; see also Palpal bulb :* Conductor: A part of the palpal bulb that accompanies and supports the embolus :* Embolus: The final part of the palpal bulb containing the end of the sperm duct, usually thin, sharp-tipped and strongly hardened (sclerotized) :* Haematodocha (plural haematodochae): A membranous, inflatable part of the palpal bulb :* Median apophysis: A projection (apophysis) of the palpal bulb, below the conductor :* Subtegulum: A hardened part of the palpal bulb nearer its base than the tegulum :* Tegulum: The main hardened part of the palpal bulb • Paracymbium: An outgrowth of the on the male • Patella: see • Paturon: The basal segment of a to which the connects • Pedicel or petiolus: The narrow connection between the and • Pedipalp (plural pedipalps or pedipalpi; also called just palp): The second appendage of the in front of the first leg; bears the in male spiders; see , see also PedipalpPlumose: Used to describe hairs () having outgrowths or appendages on two sides, giving a feather-like appearance; the appendages vary in number, size and arrangement • Pluridentate: Having multiple • Postembryo (also called larva): The stage of development between hatching from the egg and first molting • Preening brush: a dense cluster of near the tip of the posterior ; called a preening comb when present as a transverse row of setae. • Procurved: Used to describe a structure which is curved in such a way that the outer edges are in front of the central part; opposite • Prolateral: Viewed from above or below, the side of a leg or nearest the mouth, i.e. the side facing forward; opposite (includes diagram) • Promarginal: The side of the facing forward; particularly used for describing ; opposite • Prosoma: see RRastellum (plural rastella): An often rake-like structure at end of the in mygalomorph spiders; used in burrowing • Rebordered: Having a thickened edge (i.e. border) (more rarely seen as reborded, from the French , e.g. in Levy (1984)); particularly used of the • Receptaculum (plural receptacula): see • Recurved: Used to describe a structure which is curved in such a way that the outer edges are behind the central part; opposite • Retrolateral: Viewed from above or below, the side of a leg or furthest from the mouth, i.e. the side facing backwards; opposite • Retrolateral tibial apophysis: A backward-facing projection on the tibia of the male ; distinguishing feature of the RTA cladeRetromarginal: The side of the facing backward (towards the posterior end of the spider); particularly used for describing ; opposite • Rostrum (also called labrum): A component (the "upper lip") of the mouthparts, concealed by the ; see also Arthropod mouthparts: Labrum SScape: An elongated process or appendage of some • Sclerite: A single hardened (sclerotized) part of the external covering (tegument, exoskeleton) • Scopula (plural scopulae): A brush of hairs (setae); called a when on the end of the foot (tarsus), where it improves adhesion • Scutum (plural scuta): A hardened (sclerotized) plate on the of some spiders • Secondary eye: An eye belonging to the three pairs – anterior lateral eyes (ALE), posterior median eyes (PME) and posterior lateral eyes (PLE) – that are primarily movement detectors and have the light-detecting units (rhabdomeres) pointing away from the source; see , • Segments or articles of the legs and : :* Coxa (plural coxae): First leg segment, between body and trochanter; the coxa of the is heavily modified to form the or endite :* Trochanter: Second leg segment, between coxa and femur :* Femur (plural femora): Third leg segment, between trochanter and patella :* Patella (plural patellae): Fourth leg segment, between femur and tibia :* Tibia (plural tibiae): Fifth leg segment, between patella and metatarsus :* Metatarsus (plural metatarsi; also called basitarsus): Sixth leg segment, between tibia and tarsus; absent in the :* Tarsus (plural tarsi; also called telotarsus): Seventh (last) leg segment, after the metatarsus • Serrula: A row of tiny teeth along the edge of the • Seta (plural setae): A bristle; spiders have a variety of hair-like structures of increasing size that are referred to as hairs, bristles (setae) or • Sigillum (plural sigilla): A circular indentation on the outside of the spider, showing where an internal muscle is attached; particularly on the in some Mygalomorphae and on the in some AraneomorphaeSperm duct: A duct in the male used to store sperm • Spermatheca (plural spermathecae; also called receptulacum, receptulacum seminis): A structure in the of female spiders used to store sperm after insemination and before fertilization; see also SpermathecaSpigot: A small pointed or cylindrical structure at the tip of a from which silk emerges • Spine: A pointed, rigid structure on body and legs, usually with a basal joint; spiders have a variety of hair-like structures of increasing size that are referred to as hairs, bristles (setae) or spines • Spinneret: An appendage borne on the , typically one of six arranged in three pairs: anterior (anterior median, AMS), median (posterior median, PMS) and posterior (posterior lateral, PLS); silk emerges from small on the spinnerets; see also SpinneretSternum: The lower (ventral) portion of the • Stridulating organ: A series of thin ridges on a hardened part of the body; rubbing this with a matching series of short, stiff bristles (setae) elsewhere on the body creates a sound • Subadult: A spider in the last stage of development (penultimate instar) before becoming a sexually mature adult • Subtegulum: see TTapetum (plural tapeta): A light-reflecting layer in a making the eye appear pale • Tarsal claw (claw): One of a set of claws at the tip of the ; there may be a single pair, often concealed in a , or an additional third central claw, much smaller than the other two • Tarsal organ: a small pit, usually spherical and on the surface of each , believed to respond to humidityTarsus: see • Teeth: Pointed growths or bumps along the margins of the • Tegulum: see • Thoracic furrow: see • Tibia: see • Trachea (plural tracheae): A thin hardened internal tube, part of the respiratory system in many araneomorph spiders; opens on the underside of the via a tracheal spiracle; see Trachea § InvertebratesTrichobothrium (plural trichobothria): A slender hair-like structure of variable length on the legs and , arising from a special socket; used to detect air movements, including sounds; see , • Trochanter: see UUnidentate: Having a single tooth VVenter (or ventrum): The lower (ventral) portion or surface of the body or ; the adjective ventral may be applied to the lower portion or surface of any part of the body; see • Vulva: The internal copulatory organs of a female spider, including the , , and ==Abbreviations==
Abbreviations
Some abbreviations commonly found in descriptions of spider anatomy include: • ALE: anterior lateral eyes → • ALS: anterior lateral spinnerets → • AME: anterior median eyes → • DTA: dorsal tegular apophysis, on the back of the • DTiA: dorsal tibial apophysis, on the back of a • ITC: inferior tarsal claw → • LTA: lateral tegular apophysis, apophysis on the side of the • MOQ: median ocular quadrangle, the quadrangle formed by the four median eyes, → • PLE: posterior lateral eyes → • PLS: posterior lateral spinnerets → • PME: posterior median eyes → • PMS: posterior median spinnerets → • RCF: retrolateral cymbial fold, fold on the surface of the • RTA: retrolateral tibial apophysis, on the surface of a • STC: superior tarsal claw → • VTA: ventral tegular apophysis, on the underside of the • VTiA: ventral tibial apophysis, on the lower surface of a == See also ==
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