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Giant eland

The giant eland, also known as Lord Derby's eland or greater eland, is an open-forest and savanna antelope. A species of the family Bovidae and genus Taurotragus, it was described in 1847 by John Edward Gray. The giant eland is the largest species of antelope, with a body length ranging from 220–290 cm (7.2–9.5 ft). There are two subspecies: T. d. derbianus and T. d. gigas.

Etymology
The scientific name of the giant eland is Taurotragus derbianus, derived from three words: tauros, tragos, and derbianus. Tauros is Greek for a bull or bullock. Tragos is Greek for a male goat, and refers to the tuft of hair that grows in the eland's ear which resembles a goat's beard. The giant eland is also called "Lord Derby's eland" in honour of Edward Smith-Stanley, 13th Earl of Derby. It was due to his efforts that the giant eland was first introduced to England between 1835 and 1851. Lord Derby sent botanist Joseph Burke to collect animals, either alive or dead, from South Africa for his museum and menagerie. The first elands introduced in England were a pair of common elands, and what would later be identified as a giant eland bull. The details were recorded in Smith-Stanley's privately printed work, Gleanings from the Menagerie at Knowsley Hall. The Latin name indicates that it "belonged to" (given by the suffix -anus) Derby, hence derbianus. Although the giant eland is somewhat larger than the common eland, the epithet 'giant' actually refers to its large horns. ==Taxonomy==
Taxonomy
{{cladogram|align=left|title= |1= {{clade|label1= |1= {{clade |label1= |1={{clade |1={{clade |1= |2={{clade |1=Greater kudu |2={{clade |1=Mountain nyala |2={{Clade |1={{clade |1= |2={{Clade |1={{clade |1= The giant eland was first described in 1847 by John Edward Gray, a British zoologist, who called it Boselaphus derbianus. At that time, it was also called the 'black-necked eland' and Gingi-ganga. Giant eland is placed in the genus Taurotragus of family Bovidae. Giant elands are sometimes considered part of the genus Tragelaphus on the basis of molecular phylogenetics, but are usually categorized as Taurotragus, along with the common eland (T. oryx). Together with the bongo, Giant eland and common eland are the only antelopes in the tribe Tragelaphini to be given a generic name other than Tragelaphus. Although some authors, like Theodor Haltenorth, regarded the giant eland as conspecific with the common eland, they are usually considered two distinct species. ==Description==
Description
The giant elands are spiral-horned antelopes. Despite its common name, this species broadly overlaps in size with the common eland (Taurotragus oryx). However, the giant eland is somewhat larger on average than the common eland and is thus the largest species of antelope in the world. They are typically between in head-and-body length and stand approximately at the shoulder. Giant elands exhibit sexual dimorphism, as males are larger than females. The males weigh and females weigh . The tail is long, having a dark tuft of hair, and averages in length. Comparing the subspecies, T. d. derbianus is characterised by 15 body stripes, smaller size, and a rufous colour, while T. d. gigas is larger, a sandy colour, and has 12 body stripes. They can be up to long on males and on females. These features of the horns suggest that the giant eland evolved from an ancestor with true display horns. The giant eland is also parasitised by Carmyerius spatiosus (a trematode species), Taenia crocutae and T. hyaennae (two tapeworm species). ==Genetics and evolution==
Genetics and evolution
The giant eland has 31 male chromosomes and 32 female chromosomes. In a 2008 phylogenomic study of spiral-horned antelopes, chromosomal similarities were observed between cattle (Bos taurus) and eight species of spiral-horned antelopes, namely: nyala (Tragelaphus angasii), lesser kudu (T. imberbis), bongo (T. eurycerus), bushbuck (T. scriptus), greater kudu (T. strepsiceros), sitatunga (T. spekei), giant eland and common eland (T. oryx). It was found that chromosomes involved in centric fusions in these species used a complete set of cattle painting probes generated by laser microdissection. The study confirmed the presence of the chromosome translocation known as Robertsonian translocation (1;29), a widespread evolutionary marker common to all known tragelaphid species. An accidental mating between a male giant eland and a female kudu produced a male offspring, but it was azoospermic. Analysis showed that it completely lacked germ cells, which produce gametes. Still, the hybrid had a strong male scent and exhibited male behaviour. Chromosomal examination showed that chromosomes 1, 3, 5, 9, and 11 differed from the parental karyotypes. Notable mixed inherited traits were pointed ears like the eland's, but a bit widened like kudu's. The tail was half the length of that of an eland with a tuft of hair at the end as in kudu. Previous genetic studies of African savanna ungulates revealed the presence of a long-standing Pleistocene refugium in eastern and southern Africa, which also includes the giant eland. The common eland and giant eland have been estimated to have diverged about 1.6 million years ago. ==Habitat and distribution==
Habitat and distribution
Giant elands live in the broad-leafed savanna, woodlands, and glades of central and western Africa, which correspond to the two subspecies. They also live in forests as well as on the fringes of deserts. The giant elands can also live in deserts, as they produce very dry dung. They are found in South Sudan and Central African Republic into northern Cameroon and southern Chad. Recent studies proved that they also inhabit woodlands with trees of the genera Terminalia, Combretum, and Afzelia. In the past, giant elands occurred throughout the relatively narrow belt of savanna woodland that extends across West and Central Africa from Senegal to the Nile. Today they are conserved in national parks and reserves, and occur mostly in Senegal. The western giant eland is largely restricted to Niokolo-Koba National Park in Senegal. The eastern giant eland is found in several reserves, for example in Bénoué National Park, Faro National Park and Bouba Njida National Park in Cameroon and in Manovo-Gounda St. Floris National Park in the Central African Republic. They are also kept in captivity. ==Ecology and behaviour==
Ecology and behaviour
Primarily nocturnal, giant elands have large home ranges and seasonal migration patterns. They form separate groups of males and of females and juveniles. Giant elands are alert and wary, making them difficult to approach and observe or to hunt. If a bull senses danger, he will give deep-throated barks while leaving the herd, repeating the process until the whole herd is aware of the danger. Giant elands can move quickly, running at over , and despite their size are exceptional jumpers, easily clearing heights of . Diet Primarily a herbivore, the giant eland eats grasses and foliage, as well as other parts of a plant. In 2010, histological analysis of the feces of South African western giant elands was done in the Niokolo-Koba National Park and in the Bandia National Reserve. In both studies leaves, shoots of woody plants, and fruits were found to be the three major components. The other components that appeared in minor proportions were forbs and grasses, generally below five percent of the mean fecal volume. They were seen eating most foliage from Boscia angustifolia, Grewia bicolor, Hymenocardia acida, and Ziziphus mauritiana, and the fruits of Acacia and Strychnos spinosa. In the Bandia Reserve, differences in diet were marked among age classes. The conclusions were that in the dry season the eland was a pure browser, consuming grasses in small amounts. Reproduction Mating occurs throughout the year, but peaks in the wet season. Females reach sexual maturity at about two years, and males at four to five years. A female can remain in estrus for three days, and the estrous cycle is 21–26 days long. In some areas distinct breeding seasons exist. In southern Africa, females have been seen giving birth from August to October, and are joined by the males from late October to January. In Zambia calves are born in July and August. A Senegalese study focused on the suckling behaviour of giant eland and common eland calves about one to five months old determined that suckling bouts increased with the age of the calves. No other change occurred in the farmed common eland calves, but in the giant eland calves, the males were found to suckle more than female ones and shorter suckling bouts were marked in primiparous mothers than multiparous ones. The results suggest that Derby elands that lived in their natural habitat adjusted their maternal behaviour so as to be able to readily maintain a vigilant lookout for predators and other similar risks. In contrast, the farmed common elands behaved as in the conditions of captivity, without predators. ==Populations==
Populations
The eastern giant eland ranged from Nigeria, through Cameroon, Chad, the Central African Republic, and the Democratic Republic of the Congo (formerly Zaire) to Sudan and Uganda in 1980. As of 2008, a population of less than 200 individuals occur there, and only a few elands exist in neighboring countries. ==Interaction with humans==
Interaction with humans
Threats and conservation The major threats to the western giant eland population are overhunting for its rich meat and habitat destruction caused by the expansion of human and livestock populations. Its presence is uncertain in Guinea-Bissau and Nigeria. The eastern giant eland is conserved in the Faro National Park, Bénoué National Park, Bouba Njida National Park, Bamingui-Bangoran National Park and Manovo-Gounda St. Floris National Park. They are bred in captivity in the Bandia Reserve and Fathala Reserve in Senegal, Uses Giant elands give large quantities of tender meat and high-quality hides even if fed a low-quality diet. These are game animals and are also hunted for trophies. Their milk is comparatively richer in proteins and milkfat than dairy cows, which may be an explanation for the quick growth of eland calves. Eland's milk has about triple the fat content and twice the protein of a dairy cow's milk. Its docility and profitable characteristics have made it a target of domestication in Africa and Russia and has also resulted in hunting. Many people prefer to tame and raise eland rather than cattle due to their numerous benefits. Elands can survive on scarce water, which is a great advantage over domestic cattle. They can also eat coarse grasses, and can even manage to ingest some poisonous plants that can prove fatal for cattle. They are also immune to some diseases to which cattle may succumb. ==References==
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