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Frank Ruddle

Francis Hugh Ruddle (1929–2013) was an American cell and developmental biologist who was the Sterling Professor at Yale University. Ruddle was an early visionary of the Human Genome Project and created the first genetically modified mouse. He was a pioneer in both human and mouse genetics.

Early life and education
Ruddle's parents, Thomas H. Ruddle and Mary Henley Rhodda Ruddle, immigrated from the United Kingdom to West New York, New Jersey, where Frank was born on August 19, 1929. He grew up in Mariemont, Ohio, where Ruddle spent a lot of his childhood near the Ohio River. With the help of the GI Bill, Frank was able to continue his education at Wayne State University, where he earned both his bachelor's degree and master's degree just two years apart. During the year of 1960, he received his Ph.D. in zoology at the University of California at Berkeley. == Career and projects ==
Career and projects
In 1961, the Yale University's biology department welcomed Ruddle. It was here where he devoted his time to analyzing the genetics of somatic cells and recombination of DNA. Ruddle started his research and experimentation at Yale, and his work in the lab contributed to the development of the Human Genome Project. The journal documented the development of many different fields connected to gene mapping. == Awards and acknowledgements ==
Awards and acknowledgements
In 1971, Ruddle became the president of the Society for Developmental Biology, and in 1985 he was also the American Society of Human Genetics' president. Two years later, he became the president of the American Society of Cell Biology. Frank was honored many times for his achievements. He was elected to the National Academy of Sciences in 1976, the Institute of Medicine in 1985, and the American Academy of Arts and Sciences in 1977. In 1983, Ruddle received the William Alan Memorial Award by the American Society of Human Genetics. Ruddle also received the Dickson Prize in Medicine. Throughout his entire career, he contributed more than 900 publications to the field. == References ==
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