He was born on 15 September 1893, the son of Rev G C Carter and Hilda E Keane. He studied at
Marlborough College and then was awarded a place at
Cambridge University, where he continued also at postgraduate level, gaining a PhD in Zoology. His studies were interrupted by the
First World War: he served in the 6th Leicestershire Regiment from 1914-1917 and then as a Sound Ranger in the
Royal Engineers 1917 to 1919. After the war he obtained a post at the
Stazione Zoologica in
Naples where he worked 1922 to 1923 before receiving a post as a lecturer in Zoology at
Glasgow University. He stayed at Glasgow until 1930, then receiving a Fellowship from
Corpus Christi College, Cambridge, lecturing there from 1938 until retiral in 1960 . He was elected a Fellow of the
Royal Society of Edinburgh in 1925. He died in
Cambridge on 2 December 1969. ==Publications==