Golden-crowned sparrow

The golden-crowned sparrow is one of five species in the genus Zonotrichia, a group of large American sparrows. It has no subspecies. It is a sister species with, and very closely related to, the white-crowned sparrow; studies of mitochondrial DNA show the two evolved into separate species very recently in geologic time. The white-throated sparrow is a slightly more distant relative. Hybridization with both white-crowned and white-throated sparrows has been reported. Ornithologist John Latham first described the species in his book General Synopsis of Birds in 1781, but he thought it was a variety of what he called the "black-crowned bunting" and neglected to give it a scientific name. That lapse was corrected in 1789, when Johann Friedrich Gmelin assigned it the name Emberiza atricapilla in the 13th edition of Systema naturae. Various authorities in the 1800s (including John James Audubon) placed it in the genus Fringilla, but many assigned it to its current genus, Zonotrichia, once William Swainson had established that genus. Peter Simon Pallas described the same species in 1844 as Zonotrichia coronata and, for much of the 1800s, this was the name used by most authorities. In the bird's scientific name, the genus name Zonotrichia is a compound word: from Ancient Greek (, ) and (, ). The specific epithet atricapilla is Latin for "black-haired": () and (). ==Description==

Measuring in length with a wingspan of and ranging from in mass, the adult golden-crowned sparrow is fairly large for an American sparrow. Like all Zonotrichia sparrows, it has a relatively long, square-tipped tail and a slightly peaked crown. Males and females are similarly plumaged, though males average slightly larger than females. Other than a plain gray nape, the adult's upperparts are grayish-brown, with broad brownish-black streaks on the back and scapulars, and an unstreaked rump. Its underparts are gray, slightly paler on the belly and buffier on the flanks. Its wings and tail are brown, and it shows two white wing bars. Its legs are pale brown, and its bill is dark, with the upper mandible darker than the lower. Its iris is brown. In the breeding season, the golden-crowned sparrow has a broad yellow central crown stripe which becomes pale gray towards the back of the head. Similar species Although its distinctive crown patches allow for easy identification of the adult golden-crowned sparrow in breeding plumage, an immature or non-breeding bird might be mistaken for a white-crowned or white-throated sparrow. It is distinguished from the former by its crown pattern and dusky (rather than pale pink or yellowish) bill, and from the latter by its larger size, plain throat, and lack of a buff-colored central crown stripe. A young or non-breeding bird may also resemble a female house sparrow, but can be distinguished by its larger size, darker plumage, longer tail and (usually) some amount of dull yellow feathering on its forehead. ==Distribution and habitat==

The golden-crowned sparrow is common along the western edge of North America. It is a migratory species, breeding from north-central Alaska (including the Aleutian Islands as far west as Unimak Island) and central Yukon south to the northwestern corner of the US state of Washington, and wintering from southern coastal Alaska to northern Baja California. It has been recorded as a vagrant in Japan and Russia, and occasionally strays as far as the eastern coast of North America, from Nova Scotia to Florida. In the winter, it is generally found in brushy areas, (particularly chaparral), usually in dense shrubs. ==Behavior==

The golden-crowned sparrow spends its winters in flocks with conspecifics, but also often with other sparrow species, especially white-crowned sparrows. Flock territories typically range in size from . Individual birds remain with the same flock for the entire winter and return to the same wintering site each year. Breeding and lifespan The breeding season runs from late May through early August. Males on the breeding ground sing throughout the day from an exposed perch. Like other Zonotrichia sparrows, it has a sharp, distinctive call note, which is transcribed as tsew. ==Conservation and threats==

The International Union for Conservation of Nature lists the golden-crowned sparrow as a species of least concern. The sparrow's extremely large range and population size lift it well above the thresholds used to designate an imperiled species, and hippoboscid flies, including Ornithoica vicina. A number of predators prey on the golden-crowned sparrow at some point in its life cycle. These include northern and loggerhead shrikes, sharp-shinned and Cooper's hawks, northern harriers, merlins, mountain pygmy owls, western screech owls, barn owls, feral cats, and Columbian ground squirrels. ==References==