Golden jackal

The word 'jackal' appeared in the English language around 1600. It derives from the Turkish word çakal, which originates from the Persian word šagāl. The golden jackal is also known as the common jackal. ==Taxonomy==

The biological family Canidae is composed of the South American canids, the fox-like canids, and the wolf-like canids. All species within the wolf-like canids share a similar morphology and possess 78chromosomes, allowing them potentially to interbreed. Evolution }} The Arno river dog (Canis arnensis) is an extinct species of canine that was endemic to Mediterranean Europe during the Early Pleistocene around 1.9million years ago. It is described as a small jackal-like dog and probably the ancestor of modern jackals. Indian golden jackals exhibit the highest genetic diversity, and those from northern and western India are the most basal, which indicates that India was the center from which golden jackals spread. The extant golden jackal lineage commenced expanding its population in India 37,000 years ago. During the Last Glacial Maximum, 25,000 to 18,000 years ago, the warmer regions of India and Southeast Asia provided a refuge from colder surrounding areas. At the end of the Last Glacial Maximum and the beginning of the warming cycles, the golden jackal lineage expanded out of India and into Eurasia to reach the Middle East and Europe. confirming their more recent expansion into Europe. Genetic data indicates that the golden jackals of the Peloponnese Peninsula in Greece and the Dalmatian coast in Croatia may represent two ancient European populations from 6,000 years ago that have survived into modern times. Jackals were absent from most of Europe until the 19th century, when they started to expand slowly. Jackals were recorded in Hungary with the nearest population known at that time being found in Dalmatia, some 300 kilometers away. This was followed by rapid expansion of jackals towards the end of the 20th century. Golden jackals from both Southeast Europe and the Caucasus are expanding into the Baltic. In the Middle East, golden jackals from Israel have a higher genetic diversity than European jackals. This is thought to be due to Israeli jackals having hybridized with dogs, gray wolves, and African golden wolves, 13 subspecies were described since then. ==Description==

The golden jackal is similar to the gray wolf but is distinguished by its smaller size, lighter weight, more elongated torso, less-prominent forehead, shorter legs and tail, and a muzzle that is narrower and more pointed. This feature was once associated with magical powers by the people of Sri Lanka. The jackal's fur is coarse and relatively short, with the base color golden, varying seasonally from a pale creamy yellow to a dark tawny. The fur on the back is composed of a mixture of black, brown, and white hairs, sometimes giving the appearance of the dark saddle like that seen on the black-backed jackal. The underparts are a light pale ginger to cream color. Individual specimens can be distinguished by their unique light markings on the throat and chest. The coats of jackals from high elevations tend to be more buff-colored than those of their lowland counterparts while those of jackals in rocky, mountainous areas may exhibit a grayer shade. The bushy tail has a tan to black tip. Melanism can cause a dark-colored coat in some golden jackals, a coloring once fairly common in Bengal. Unlike melanistic wolves and coyotes that received their dark pigmentation from interbreeding with domestic dogs, melanism in golden jackals probably stems from an independent mutation that could be an adaptive trait. What is possibly an albino specimen was photographed in southeastern Iran during 2012. The jackal moults twice a year, in spring and in autumn. In Transcaucasia and Tajikistan, the spring moult begins at the end of winter. If the winter has been warm, the spring moult starts in the middle of February; if the winter has been cold, it begins in the middle of March. The spring moult lasts for 60–65 days; if the animal is sick, it loses only half of its winter fur. The spring moult commences with the head and limbs, extends to the flanks, chest, belly and rump, and ends at the tail. Fur on the underparts is absent. The autumn moult occurs from mid-September with the growth of winter fur; the shedding of the summer fur occurs at the same time. The development of the autumn coat starts with the rump and tail and spreads to the back, flanks, belly, chest, limbs and head, with full winter fur being attained at the end of November. ==Ecology==

The golden jackal inhabits Europe and Southwest, Central, South, and Southeast Asia. In Finland, a golden jackal has been trapped near Sodankylä within the Arctic Circle. Diet carcass in Pench National Park The golden jackal fills much the same ecological niche in Eurasia as the coyote does in North America; it is both a predator and a scavenger, Competition and its kits in south-western Germany The jackal's competitors are the red fox, wolf, jungle cat, wildcat, and raccoon in the Caucasus, and the steppe wildcat in Central Asia. Red foxes and golden jackals share similar diets. Red foxes fear jackals, which are three times bigger than them. Red foxes will avoid close proximity to jackals and fox populations decrease where jackals are abundant. Striped hyenas prey on golden jackals, and three jackal carcasses were found in one hyena den. ==Behavior==

Social behavior Golden jackals exhibit flexible social organization depending on the availability of food. The breeding pair is the basic social unit, and they are sometimes accompanied by their current litter of pups. In India, their distributions are a single jackal, 31%, two jackals, 35%, three jackals, 14%, and more than three jackals, 20%. Family groups of up to 4–5 individuals have been recorded. Scent marking through urination and defecation is common around golden jackal den areas and on the trails they most often use. Scent marking is thought to assist in territorial defense. The hunting ranges of several jackals can overlap. Jackals can travel up to during a single night in search of either food or more suitable habitat. Non-breeding members of a pack may stay near a distant food source, such as a carcass, for up to several days before returning to their home range. Home range sizes can vary between , depending on the available food. {{CSS image crop Reproduction Golden jackals are monogamous and will remain with the one partner until death. Female jackals have only one breeding cycle each year. Breeding occurs from October to March in Israel and from February to March in India, Turkmenistan, Bulgaria, and Transcaucasia, with the mating period lasting up to 26–28 days. Females undergoing their first estrus are often pursued by several males that may quarrel among themselves. Mating results in a copulatory tie that lasts for several minutes, as it does with all other canids. Gestation lasts 63 days, and the timing of the births coincides with the annual abundance of food. In India, the golden jackal will take over the dens of the Bengal fox and the Indian crested porcupine, and will use abandoned gray wolf dens. Most breeding pairs are spaced well apart and maintain a core territory around their dens. Den excavations commence from late April to May in India, with dens located in scrub areas. Rivulets, gullies, and road and check-dam embankments are prime denning habitats. Drainage pipes and culverts have been used as dens. Dens are long and deep, with between 1–3 openings. Young pups can be moved between 2–4 dens. The male helps with digging the den and raising the pups. In the Caucasus and Transcaucasia, the burrow is located either in thick shrub, on the slopes of gullies, or on flat surfaces. In Dagestan and Azerbaijan, litters are sometimes located within the hollows of fallen trees, among tree roots, and under stones on river banks. In Central Asia, the golden jackal does not dig burrows but constructs lairs in dense tugai thickets. Jackals in the tugais and cultivated lands of Tajikistan construct lairs in long grass, shrubs, and reed openings., IsraelIn Transcaucasia, golden jackal pups are born from late March to late April, and in northeastern Italy during late April; they can be born at any time of year in Nepal. The number of pups born in a single litter varies geographically. Jackals in Transcaucasia give birth to 3–8 pups, Tajikistan 3–7 pups, Uzbekistan 2–8 pups, and Bulgaria 4–7 pups; in India the average is four pups. The pups are born with closed eyes that open after 8–11 days, with the ears erecting after 10–13 days. Their teeth erupt at 11 days after birth, and the eruption of adult dentition is completed after five months. Pups are born with soft fur that ranges in color from light gray to dark brown. At the age of one month, their fur is shed and replaced with a new reddish-colored pelt with black speckles. The pups have a fast growth rate and weigh at two days of age, at one month, and at four months. Females possess four pairs of teats, and lactation lasts for up to 8–10 weeks. The pups begin to eat meat at the age of 15–20 days. Dog pups show unrestrained fighting with their siblings from 2 weeks of age, with injury avoided only due to their undeveloped jaw muscles. This fighting gives way to play-chasing with the development of running skills at 4–5 weeks. Wolf pups possess more-developed jaw muscles from 2 weeks of age, when they first show signs of play-fighting with their siblings; serious fighting occurs during 4–6 weeks of age. Compared to wolf and dog pups, golden jackal pups develop aggression at the age of 4–6 weeks, when play-fighting frequently escalates into uninhibited biting intended to harm. This aggression ceases by 10–12 weeks when a hierarchy has formed. Once the lactation period concludes, the female drives off the pups. Pups born late remain with their mother until early autumn, at which time they leave either singly or in groups of two to four individuals. Females reach sexual maturity after 10–11 months and males at 21–22 months. Foraging The golden jackal often hunts alone, and sometimes in pairs, but rarely hunts in a pack. When hunting alone, it trots around an area and occasionally stops to sniff and listen. Once prey is located, the jackal conceals itself, quickly approaches its prey and then pounces on it. Single jackals hunt rodents, hares, and birds. They hunt rodents in grass by locating them with their hearing before leaping into the air and pouncing on them. In India, they can dig Indian gerbils out from their burrows, and they can hunt young, old, and infirm ungulates up to 4–5 times their body weight. Jackals search for hiding blackbuck calves throughout the day during the calving period. The peak times for their searches are the early morning and the late evening. When hunting in pairs or packs, jackals run parallel to their prey and overtake it in unison. When hunting aquatic rodents or birds, they will run along both sides of narrow rivers or streams and drive their prey from one jackal to another. Pack-hunting of langurs is recorded in India. Packs of between 5 and 18 jackals scavenging on the carcasses of large ungulates is recorded in India and Israel. Packs of 8–12 jackals consisting of more than one family have been observed in the summer periods in Transcaucasia. In India, the Montagu's harrier and the Pallid harrier roost in their hundreds in grasslands during their winter migration. Jackals stalk close to these roosting harriers and then rush at them, attempting to catch one before the harriers can take off or gain sufficient height to escape. In Southeastern Asia, golden jackals have been known to hunt alongside dhole packs. They have been observed in the Blackbuck National Park, Velavadar, India, following Indian wolves (Canis lupus pallipes) when these are on a hunt, and they will scavenge off wolf kills without any hostility shown from the wolves. In India, lone jackals expelled from their pack have been known to form commensal relationships with tigers. These solitary jackals, known as kol-bahl, will associate themselves with a particular tiger, trailing it at a safe distance to feed on the big cat's kills. A kol-bahl will even alert a tiger to prey with a loud "pheal". Tigers have been known to tolerate these jackals, with one report describing how a jackal confidently walked in and out between three tigers walking together. Golden jackals and wild boar can occupy the same territory. ==Conservation==

, India , Odisha, India. The golden jackal is listed as Least Concern on the IUCN Red List due to its widespread distribution, with it being common throughout its range and with high densities in those areas where food and shelter are abundant. In Europe, golden jackals are not listed under the 1973 Convention on International Trade in Endangered Species of Wild Fauna and Flora nor the 1979 Convention on the Conservation of Migratory Species of Wild Animals. Golden jackals in Europe fall under various international legal instruments. These include the 1979 Berne Convention on the Conservation of European Wildlife and Natural Habitats, the 1992 Convention on Biological Diversity, and the 1992 European Union Council Directive 92/43/EEC on the Conservation of Natural Habitats and of Wild Fauna and Flora. The Council Directive provides both guidance and limits on what participating governments can do when responding to the arrival of expanding jackals. These legislative instruments aim to contribute to conserving native wildlife; some governments argue that the golden jackal is not native wildlife but an invading species. The Golden Jackal informal study Group in Europe (GOJAGE) is an organization that is formed by researchers from across Europe to collect and share information on the golden jackal in Europe. The group also has an interest in the golden jackal's relationship with its environment across Eurasia. Membership is open to anyone who has an interest in golden jackals. In Europe, there are an estimated 70,000 golden jackals. They are fully protected in Albania, North Macedonia, Germany, Italy, Poland and Switzerland. They are unprotected in Belarus, Bosnia and Herzegovina, Czech Republic, Estonia, and Greece. They are hunted in Bosnia and Herzegovina, Bulgaria, Croatia, Hungary, Kosovo, Latvia, Lithuania, Montenegro, Romania, Serbia, Slovakia, Slovenia, and Ukraine. Their protection in Austria and Turkey depends on the part of the country. Their status in Moldova is not known. The Syrian jackal was common in Israel and Lebanon in the 1930s–40s, but their populations were reduced during an anti-rabies campaign. Its current status is difficult to ascertain, due to possible hybridisation with pariah dogs and African golden wolves. The jackal population for the Indian subcontinent is estimated to be over 80,000. In India, the golden jackal occurs in all of India's protected areas apart from those in the higher areas of the Himalayas. It is included in CITES AppendixIII, and is listed in the Wildlife Protection Act, 1972, under ScheduleIII, thus receiving legal protection at the lowest level to help control the trade of pelts and tails in India. ==Relationships with humans==

In folklore, mythology and literature (left) torments Father Wolf and his family, as illustrated in the 1895 edition of Rudyard Kipling's The Two Jungle Books. Golden jackals appear in Indian folklore and in two ancient texts, the Jakatas and the Panchatantra, where they are portrayed as intelligent and wily creatures. One popular Indian saying describes the jackal as "the sharpest among beasts, the crow among birds, and the barber among men". For a person embarking on an early morning journey, hearing a jackal howl was considered to be a sign of impending good fortune, as was seeing a jackal crossing a road from the left side. The high predation rate by jackals in both Bulgaria and Israel is attributable to the lack of preventative measures in those countries and the availability of food in illegal garbage dumps, leading to jackal population explosions. Golden jackals are extremely harmful to fur-bearing rodents, such as coypu and muskrats. Coypu can be completely extirpated in shallow water bodies. During 1948–1949 in the Amu Darya, muskrats constituted 12.3% of jackal fecal contents, and 71% of muskrat houses were destroyed by jackals. Jackals also harm the fur industry by eating muskrats caught in traps or taking skins left out to dry. Hunting During British rule in India, sportsmen conducted golden jackal hunting on horseback with hounds, with jackal coursing a substitute for the fox hunting of their native England. They were not considered as beautiful as English red foxes, but were esteemed for their endurance in the chase with one pursuit lasting hours. India's weather and terrain added further challenges to jackal hunters that were not present in England: the hounds of India were rarely in as good condition as English hounds, and although the golden jackal has a strong odor, the terrain of northern India was not good in retaining scent. Also, unlike foxes, jackals sometimes feigned death when caught and could be ferociously protective of their captured packmates. Jackals were hunted in three ways: with greyhounds, with foxhounds, and with mixed packs. Hunting jackals with greyhounds offered poor sport because greyhounds were too fast for jackals, and mixed packs were too difficult to control. From 1946 in Iraq, British diplomats and Iraqi riders conducted jackal coursing together. They distinguished three types of jackal: the "city scavenger", which was described as being slow and so smelly that dogs did not like to follow them; the "village jack", which was described as being faster, more alert, and less odorous; and the "open-country jack", which was described as being the fastest, cleanest, and providing the best sport of all three populations. Some indigenous people of India, such as the Kolis and Vagris of Gujarat and Rajasthan and the Narikuravas in Tamil Nadu, hunt and eat golden jackals, but the majority of South Asian cultures consider the animal to be unclean. The orthodox dharma texts forbid the eating of jackals because they have five nails. In the area of the former Soviet Union, jackals are not actively hunted and are usually captured only incidentally during the hunting of other animals by means of traps or shooting during drives. In Transcaucasia, jackals are captured with large fishing hooks baited with meat and suspended from the ground with wire. The jackals can only reach the meat by jumping, and are then hooked by the lip or jaw. Fur use In Russia and the other nations of the former Soviet Union, golden jackals are considered furbearers of low quality because of their sparse, coarse, and monotonously colored fur. Jackal hairs have very little fur fiber; therefore, their pelts have a flat appearance. The jackals of Asia and the Middle East produce the coarsest pelts, though this can be remedied during the dressing process. Elburz in northern Iran produces the softest furs. Jackal skins are not graded to a fur standard, and are made into collars, women's coats, and fur coats. During the 1880s, 200 jackals were captured annually in Mervsk and in the Zakatal area of the Transcaucasus, with 300 jackals being captured there during 1896. In this same period, a total of 10,000 jackals were taken within Russia and their furs sent exclusively to the Nizhegorod fair. In the early 1930s there were 20,000–25,000 jackal skins tanned annually in the Soviet Union, but these could not be utilized within the country, and so the majority were exported to the United States. Commencing from 1949, they were all used within the Soviet Union. Sulimov dog , Russia The golden jackal may have once been tamed in Neolithic Turkey 11,000 years ago, as there is a sculpture of a man cradling a jackal found in Göbekli Tepe. French explorers during the 19thcentury noted that people in the Levant kept golden jackals in their homes. The Kalmyk people near the Caspian Sea were known to frequently cross their dogs with jackals, and Balkan shepherds once crossed their sheepdogs with jackals. The Russian military established the Red Star kennels in 1924 to improve the performance of working dogs and to conduct military dog research. The Red Star kennel developed "Laikoid" dogs, which were a cross-breed of Spitz-type Russian Laikas with German Shepherds. By the 1980s, the ability of Russia's bomb and narcotic detection dogs were assessed as being inadequate. Klim Sulimov, a research scientist with the DS Likhachev Scientific Research Institute for Cultural Heritage and Environmental Protection, began cross-breeding dogs with their wild relatives in an attempt to improve their scent-detection abilities. The researchers assumed that during domestication dogs had lost some of their scent-detection ability because they no longer had to detect prey. Sulimov crossed European jackals with Laikas, and also with fox terriers to add trainability and loyalty to the mix. He used the jackal because he believed that it was the wild ancestor of the dog, that it had superior scent-detecting ability, and, because it was smaller with more endurance than the dog, it could be housed outdoors in the Russian climate. Sulimov favored a mix of one quarter jackal and three-quarters dog. Sulimov's program continues today with the use of the hybrid Sulimov dogs at the Sheremetyevo Airport near Moscow by the Russian airline Aeroflot. The hybrid program has been criticized, with one of Sulimov's colleagues pointing out that in other tests the Laika performed just as well as the jackal hybrids. The assumption that dogs have lost some of their scent-detection ability may be incorrect, in that dogs need to be able to scent-detect and identify the many humans that they come into contact with in their domesticated environment. Another researcher crossed German Shepherds with wolves and claimed that this hybrid had superior scent-detection abilities. The scientific evidence to support the claims of hybrid researchers is minimal, and more research has been called for. == Notes ==