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Eastern woodrat

The eastern woodrat, also known as the Florida woodrat or bush rat, is a pack rat native to the central and Eastern United States. It constructs large dens that may serve as nests for many generations and stores food in outlying caches for the winter. While widespread and not uncommon, it has declined or disappeared in several areas.

Taxonomy
Eight subspecies of the eastern woodrat are currently recognized: N. f. illinoensis, N. f. floridana (the nominate), N. f. smalli, N. f. baileyi, N. f. pennsylvanica, N. f. attwateri, N. f. osagensis, and N. f. rubida. Of these, the Key Largo woodrat (N. f. smalli) is classified as endangered by the IUCN. The Allegheny woodrat (N. magister) was previously considered a subspecies but was elevated to species status based on comparative mitochondrial DNA analyses. Subspecies are thought to be descended from one species living along the Appalachian Plateau, which subsequently spread out through the Coastal Plain. == Description ==
Description
The eastern woodrat is a rodent of medium size, with an average length of and weight of , The body is short and stocky and the tail is exceptionally long (). It is covered in a soft, gray-brown pelage, which is darker dorsally. Belly and feet are white. Sexes are alike, but males tend to be slightly larger. == Distribution and habitat ==
Distribution and habitat
Distribution The distribution for the eastern woodrat stretches across the Southeastern and Midwestern United States. They are located as far south as the Tennessee River and Central Florida. The more central areas they are found are Kentucky and Tennessee. Northern locations include Kansas, central Missouri, and southern Illinois. They can also be found along the Appalachian Mountains that range all the way to New York. They can be seen in places like woodlands, prairies, mountains, swamps, and lowland hardwood forests as far west as Colorado. The more disjunctive population occurrence is in Nebraska and Key Largo, Florida. With a wide range but low population density, this species is considered uncommon. The overall population has been decreasing since 1982. The primary cause for this decline in the southeastern United States has been habitat destruction due to human development of the coastal plains. The species has been recovered as a fossil from late Pleistocene deposits in southeastern New Mexico, several hundred miles southwest of its nearest current range. The species does not hibernate, even in the northern extremes of its range. Habitat The species inhabits wooded areas, swamps and hedges, with a variety of different habitat preferences reported for the recognized geographic subspecies. Eastern woodrats are typically solitary creatures, so they usually have a buffer between territories. The females have a smaller territory than males, averaging around . Males average around and the species-wide average territory area is . Being solitary, the rats tend not to stray much farther than from their nests unless sexually active or for a preferred food. ==Ecology==
Ecology
Foraging As with most members of the genus, the Eastern woodrat feeds opportunistically on nuts, seeds, fungi, buds, stems, roots, foliage, and fruits. While the eastern woodrat's nest is typically found on the ground, it is a capable climber and may forage above ground. Eastern woodrats eat about 5% of their body weight in dry mass each day. During the summer months, most feeding is done while foraging. Only small amounts of food are taken back to the den for daytime feeding. Eastern woodrats are known for their foraging and caching habits. When searching for food is dangerous or unproductive, animals often use food stores to supply all or part of their diet. This is a feasible strategy to avoid food shortage. It is the habit of collecting and storing both food and nonfood items that has earned the eastern woodrat is other common name of "pack-rat" or "trade rat". Starting in September, the woodrat will start to forage and store food in its midden for use and survival in the winter. Although caches do not serve as the sole source of winter food, caches examined yielded as much as of plant material. Decomposition of food stores appears to be a constant challenge. Woodrats seem to exhibit physiological adaptions that allow them to consume food inhabited by fungi. Fungi can increase nutritional value of some foods by making nutrients within food more accessible by breaking down complex carbohydrates, which woodrats may exploit. Although eastern woodrats are herbivores, evidence suggests that the species will take meat if given the opportunity. Snakes, salamanders, mice, and quail have all been found in stomachs of woodrats. Gnawed bones have been found in caches, probably used for sharpening teeth and for their mineral contents. Only anecdotal evidence exists concerning woodrat carnivory. However, woodrats will cache carrion if given the chance. The breeding season of eastern wood rats depends on the climate. Those in warmer climates (e.g. Florida and Georgia) can reproduce all year, while eastern wood rats in higher latitudes (e.g. Kansas and Nebraska) breed from early spring to mid fall. Their estrous cycle lasts between 3 and 8 days, while gestation lasts between 32 and 38 days. On the 15th day their hair is fully grown and their eyes are wide open. The young wean for 3–4 weeks and become independent after 70–90 days. Juveniles continue growing until they are about 8 months old. In captivity the eastern woodrat has been recorded to live for up to 8.6 years, however the average lifespan in the wild is 3 years. The majority of deaths occur within their first year of life. One field study in Kansas tracked 27 individuals, of which six survived to adulthood and only three lived long enough to reproduce. One of the most common parasites of eastern woodrats are botfly larvae. Adult botflies lay their eggs outside the entrance of the woodrat's den. They then attach themselves to the woodrat's fur when it passes through entrance. Once the eggs hatch, the botfly larvae penetrate the skin and lodge in the woodrat's neck, chest, and abdomen until pupation. The resulting cyst can be 15 mm in diameter but does not seem to cause any obvious discomfort. Botflies infest approximately 16% of the eastern woodrat population. Raccoons may carry raccoon roundworms, an intestinal parasite. Woodrats may ingest the eggs of the roundworm while foraging at raccoon latrine areas. Larvae migrate to the brain, causing a lack of energy, loss of muscle control, and eventually death. The roundworm is a known mortality factor in woodrats in Indiana, New York, New Jersey, and Pennsylvania, with infection rates of around 75%. == Ecological role ==
Ecological role
Eastern woodrat nests provide homes for a number of other species including, but not limited to, other rodents, insects, reptiles, and frogs. Seed dispersal by caching and transporting seeds into dens has a great impact on the spread and maintenance of forest ecosystems, and woodrat fecal matter increases soil fertility. The study of feces from prehistoric woodrat middens has been of great use in archaeological and paleontological research by providing indications about changing floral regimes. == Interaction with humans ==
Interaction with humans
Eastern woodrats are popularly known for being pests. They tend to seek out houses, especially cottages or cabins in wooded areas, for warmth or housing, and do not hesitate to make nests out of mattresses and other furniture while feeding in adjacent cultivated areas. Automobiles may be damaged by chewing on wires and the introduction of nesting materials. Rarely, eastern woodrats may be the carriers of diseases that are communicable to humans. The eastern woodrat does not have any economic impact on humans, but may receive blame for damages made by black or Norway rats. == Conservation ==
Conservation
Most subspecies of the eastern woodrat have been classified as Least Concern by the IUCN, with the exception of the Key Largo woodrat. While the species appears to never have been abundant, it remains widespread and reasonably common. While woodrats forage on many different food items, they are largely depended on cached acorns and nuts during winter. Eruptions of the gypsy moth in the basin of the Lower Mississippi River in 1964 and 1965 resulted in poor acorn and chestnut crops and subsequently in increased mortality in eastern woodrats in the area. Another step towards preserving this species is the separation of human activities from woodrat habitats to reduce human disturbance and secondary effects such as the presence of raccoons. == References ==
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