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Graphis subintermedians

Graphis subintermedians is a species of script lichen in the family Graphidaceae. This bark-dwelling lichen forms white to pale grey crusts with short, sparsely branched fruiting structures that have finely lined edges and produce large, multi-chambered spores. It is known only from the rainforests of Sabah in Malaysian Borneo, where it was originally collected by the American lichenologist Mason Hale.

Taxonomy
The species was formally described in 2009 by Robert Lücking, following Mason Hale's earlier unpublished manuscript name. The type locality is Sabah, Malaysian Borneo, where the holotype specimen was collected by Hale and is housed at the United States National Herbarium (US). Graphis subintermedians is placed in a small group of taxa characterised by a laterally (blackened) , striate (finely lined) , a clear hymenium, and (multi-chambered) ascospores. It differs from the otherwise similar Graphis pseudoserpens in having larger ascospores, and within this group it is the only species that both lacks detectable secondary metabolites and has ascospores exceeding 60 × 16 μm. ==Description==
Description
The thallus is (possessing a distinct outer ), smooth to uneven in texture, and white to pale grey in colour. The are to prominent, have a basal (rim of thallus tissue at the base), and are short and sparsely branched, measuring 1–3 mm long by 0.3–0.4 mm wide. Their edges () are striate. The excipulum is laterally carbonised, and the hymenium is clear (lacking oil droplets or ). Each ascus contains two large ascospores, which are hyaline, muriform, and measure 40–75 × 20–30 μm. ==Habitat and distribution==
Habitat and distribution
No lichen secondary metabolites have been detected in this species. Graphis subintermedians is known from Sabah in Malaysian Borneo, where it grows on bark in forested habitats. ==See also==
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