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Helicodiceros

Helicodiceros muscivorus, the dead horse arum lily, is an ornamental plant native to Corsica, Sardinia and the Balearic Islands. It is the only species in the genus Helicodiceros. Within the family Araceae the plant is part of the subfamily Aroideae.

Description
The inflorescence of the arum lilies is a three-part spadix which resembles the anal area of a dead mammal. In between is a hairy spathe such as a 'tail' running down into the chamber of the flower which bonds with the fertile male and female florets. The appendix and the male florets are thermogenic, but have different temporal patterns. The exits of the female florets are hindered by spines and filaments which serve to trap the blow flies once inside. The male florets exhibit independence from the ambient temperatures as heat production depends on the time of the day rather than ambient temperatures. Also, uncoupling protein was found in both the thermionic male florets and the appendix. The protein is 1178 nucleotide in length in the dead horse arum mRNA excluding the poly-A tail and it is believed to have a protein of 304 amino acids. It also possesses three mitochondrial carrier signature domains, six membrane-spanning domains, and one nucleotide-binding domain. Potato and rice have been compared to the plant at times due to its typical features of the uncoupling protein. Uncoupling protein plays a role in the production of energy to become heat. ==Thermogeny==
Thermogeny
The dead horse arum manipulates the heat to release an odor that lures the flies to the structure of the appendix of the flower to begin pollination. This odor is a strong, putrid smell, its composition bearing similarity to a real carcass, and which flies are unable to distinguish from a real carcass. Blow flies, such as the common green bottle fly (Lucilia sericata), find the smell and the flesh-colored hairy inflorescence of the plant irresistible, such that large numbers of flies are attracted into the plant.(This source should not be cited to describe the specific interaction between these two species) The thermogeny has a direct effect on the pollinators, by altering their behavior. Although male florets of the dead horse arum exhibit some independence from ambient temperature the pattern has shown that heat production depends on time of the day. Pollination occurs in one to two days. The highest temperature of the plant was found to peak at noon on day 1. The appendix temperature was 12.4 °C higher than ambient temperature. During day 1, eight flowering plants received a total of 881 fly visits. Thermogeny has been linked to be produced by the uncoupling protein; it does not complete the process to ATP synthesis, instead it allows electrons to flow into the mitochondria without creating ATP. The result is the creation of more energy which then dissipates into heat. ==Pollination==
Pollination
The dead horse arum has a two-day process for pollination. The individual flower is able to receive pollen for one day only, and usually that day its male parts are not mature. Although the male part is able to produce pollen the next day, the female part shrivels up and cannot receive it. Both these mechanisms favor pollination from another plant and discourage self-pollination. Some blowflies, such as Calliphora vomitoria, are known pollinators. ==Gallery==
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