Heracles inexpectatus

The description of a new species and genus was published in 2019, emerging from research undertaken by Trevor Worthy, Suzanne Hand, Mike Archer, R. Paul Scofield and Vanesa L. De Pietri. Alan Tennyson's and Te Papa's key role in this discovery was later acknowledged in a correction published in 2021. Heracles inexpectatus was described from two incomplete tibiotarsi (leg bones), a left one and a right one, probably from the same individual, that were discovered in 2008. The left bone was nominated as the holotype. The fossils were originally presumed to be from an eagle, but were re-inspected after a PhD student at Flinders University noted that the bone morphology was significantly different from typical eagles. The etymology of the generic epithet Heracles alludes to the earlier naming of a genus, Nelepsittacus, also found in the St Bathan Fauna, which referred to a mythical king Neleus who was slain by the Greek hero Herakles, who also slew all of his sons, save for Nestor. The authors distinguished the new species with the epithet inexpectatus to describe their astonishment at the unexpected discovery of an enormous parrot-like bird from the Miocene epoch. The etymology of the nickname "Squawkzilla" is a portmanteau of the words "squawk", an onomatopoeic word for the sound of a parrot, and "-zilla", a suffix denoting large size, derived from the fictional giant creature Godzilla. == Description ==

The largest known species of Psittaciformes, which comprises the modern parrots and cockatoos, it is estimated to have been around one meter in height, with a body mass of seven kilograms, and presumed to have been flightless, terrestrial and perhaps arboreal. Island gigantism has been observed in other orders of birds, especially in New Zealand and Fiji, but this species exceeds the proportions of any extant or fossil species of the parrot order. The previously known record for size was the arboreal and nocturnal Strigops habroptilus, the kākāpō of modern New Zealand. The fossilised tarsi were deposited in a rich and mixed assemblage of animal remains, including other large species of Aves such as the moa, anatids and an eagle, the bones of which are usually fragmented. The type material of Heracles inexpectatus is the longest evidence obtained at the site, which rarely produces fossil longer than 100 millimeters. The time of deposition is determined as sixteen to nineteen million years ago, in an area associated with a freshwater lacustrine system, in wet forest dominated by cycad, palm and casuarina species, along with a diverse array of other avian fauna. The species appears to have occupied a niche in the region's ecology, unfilled by mammals, that favoured the insular gigantism exhibited by some other avian species of the region. == Environment ==

The climate in New Zealand was similar to nowadays (temperate) and may have been subject to periodic droughts. Summer season (December to February) had average daytime temperatures between and a humidity around 80%. Winter season (June to August) had average daytime temperatures ranging from and a humidity around 67%. == Paleontology ==

The tibia fossils were discovered in a trench excavation of the foot of hills 50 meters east from the river bank of the Manuherikia River in Home Hills Station, Otago, NZ. These fossils were collected on 14 January in 2008. == References ==