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Hirticlavula

Hirticlavula is a fungal genus in the family Clavariaceae. It contains a single described species, Hirticlavula elegans. Formally described in 2014, the fungus has been collected from Norway and Denmark. H. elegans produces white fruit bodies up to 1.1 millimetres (0.043 in) in height. Each fruit body contains a fertile head atop a hairy stem. The fruit bodies grow directly from dead bark or wood, where they feed as saprotrophs. Both morphological and ecological details are distinctive when compared to other clavarioid fungi.

Taxonomy
Hirticlavula elegans was described based on collections dating back to 1995. It was provisionally reported in 2009 in the proceedings of a 2008 conference, before being formally described in 2009 by Jens H. Petersen and Thomas Læssøe in an article in Karstenia, coauthored with Marie L. Davey. The specific name elegans is from the Latin meaning "elegant", and refers to how "pretty" the species's fruit bodies are. ==Description==
Description
Hirticlavula elegans produces white fruit bodies from 800 to 1100 micrometres (μm) in height. The robust and fertile head measures from 230 to 260 by 90 to 200 μm, while the hairy stem measures between 600 and 800 by 40 to 60 μm. The fruit bodies mostly retain their shape when dried, but do develop a cream tinge. The fruit bodies do not emerge from a sclerotium, but instead grow directly from the substrate. The basidia are cyanophilous, meaning that the cell walls will readily absorb methyl blue stain. They measure 14 to 22 by 3.5 to 5 μm, and each sport four "curved and remarkably thin" sterigmata. The sterigmata are 4–5 μm in length. The smooth, broadly ellipsoid spores measure 4.5 to 6.5 by 3–4.2 μm. They are hyaline, and either cyanophilous or containing cyanophilous particles. They contain no prominent crystals, and are topped by a 0.8 μm-long apiculus (the section that connects to the sterigmatum). The hyphae lack clamp connections, and the hyphal structure is monomitic, meaning that only generative hyphae are present. In the stem, the 1–2 μm-wide hyphae run parallel to one another, but are less regular towards the base. Similar species Hirticlavula elegans fruit bodies are highly distinctive. The characteristic hairy stems are very different from those of any other species in the Clavariaceae. Hairy stems are seen among more distant relatives, including members of Typhula, Pistillina and Pterula. However, the hairs of H. elegans are specialised structures containing numerous cells, while the more irregular hairs on the stems of the other species are merely branched hyphae. In addition, the hairs on the H. elegans stems thicken at the furthest point from the stem, while any hairs on the stems of other known species of clavarioid fungi thin at the end. ==Distribution and ecology==
Distribution and ecology
Hirticlavula elegans is known from Denmark and Norway. The fruit bodies grow directly from the wood or, more usually, the bark of hardwoods. Identified substrates include oaks, willows, hazels and possibly birches. ==See also==
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