The
monophyly of the family is not in doubt. Regarding the relationships among the species of
Hygrobia, a recent phylogenetic analysis suggested a sister group relationship between
H. hermanni and a clade formed by the Australian species, with
H. nigra sister to
H. australasiae. Hygrobiidae probably diverged from other Hydradephagan clades around the time of the initial breakup of
Pangea. The split between today's
Palearctic and Australian clades occurred later, possibly in the middle
Mesozoic, by dispersal events. == Species diversity and distribution ==