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Hyperailurictis

Hyperailurictis is an extinct genus of felid from Miocene North America. The Hyperailurictis species are Pseudaelurus-grade felids and thought to be the first felids in the Americas.

Taxonomy and evolution
Taxonomic history In 1858, the paleontologist Joseph Leidy described a new species of cat, Felis intrepidus, based on lower jaw fragments (a left ramus, described, and a right ramus, mentioned only and now lost) found somewhere in Nebraska near the Niobrara river. The fragments were later determined to have come from the lower Valentine Formation, making them late Barstovian in age. Leidy reassigned the specimens to Pseudaelurus as Pseudaelurus intrepidus in a 1869 paper. In 1873, an expedition collected another lower jaw from the lower Valentine Formation, which was described by Thorpe in 1922 as the holotype of the new species Pseudaelurus marshi. He also described a paratype, a left ramus collected from part of the Ogallala Group (also of Barstovian age). Shortly after, he also described Pseudaelurus thinobates, a larger species from a late Clarendonian locality in California. However, David Kitt removed Pseudaelurus thinobates from the genus, placing it instead as Nimravides thinobates, during a 1958 study. Subsequently, a review of the genus reassigned Pseudaelurus pedionomus to the new genus as Nimravides pedionomus. The fragment remained the only specimen assigned to Ps. aeluroides, and an examination of illustration and measurements of the fragment in 2003 lead Tom Rothwell to suggest that it actually belonged to a juvenile Ps. marshi. Similar in size to Nimravides pedionomus, this was referred in 2003 to Nimravides hibbardi instead. He also referred a number of other specimens, collected from early Barstovian localities in Nebraska and New Mexico and from late Barstovian localities in New Mexico and Colorado, to Pseudaelurus stouti. However, this was largely ignored by later studies until a 2010 review of the Felidae as a whole brought up the proposal again, leading to some authors, though not all, recognizing the generic split. ==Description==
Description
Hyperailurictis intrepidus was a relatively large species, comparable in size to a modern-day leopard, and overlapping in size with the Eurasian Pseudaelurus quadridentatus. H. marshi was also a large species, but differed from H. intrepidus because of certain mandibular features. H. stouti, which was contemporary with the two, was by contrast a very small and slender felid. H. validus overlaps in size with H. intrepidus and H. marshi, being similar in size to a leopard. The final species, H. skinneri is much smaller than H. validus, overlapping in size with the Eurasian species Pseudaelurus lorteti, P. cuspidatus, and P. guangheensis. ==Distribution and habitat==
Distribution and habitat
Hyperailurictis validus and Hyperailurictis skinneri are the earliest recorded species, both occurring in various late Hemingfordian localities; H. validus in New Mexico and Nebraska, and H. skinneri only in Nebraska. H. validus is also known from several early Barstovian localities in Nebraska. Also known from early Barstovian-aged localities are the species H. intrepidus (localities in Texas, Colorado, Nevada, California, and Nebraska), H marshi (localities in Texas, California, Colorado, and Nebraska) and H. stouti (from localities in Nebraska and New Mexico). The dubious species H. aeluroides is also based on a fossil from the early Barstovian; the fossil in question was found in Nebraska. In addition, H. intrepidus, H. marshi, and H. stouti are all known from localities of late Barstovian age. H. intrepidus and H. marshi are both known from localities in Nebraska and California, with H. marshi additionally occurring in localities in Colorado and New Mexico. H. stouti has been recorded in localities in Colorado and New Mexico as well. ==References==
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