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Kutorginata

Kutorginates (Kutorginata) are an extinct class of early rhynchonelliform ("articulate") brachiopods. The class contains only a single order, Kutorginida (kutorginides). Kutorginides were among the earliest rhynchonelliforms, restricted to the lower-middle part of the Cambrian Period.

Anatomy
Kutorginides typically have a ventribiconvex shell (both valves convex, the ventral valve moreso) and a strophic (straight) hinge line. Based on fossils of Nisusia, the shell’s internal (secondary) layer appears to have a microstructure of calcite fibers. Several kutorginides are preserved in lagerstätten, elaborating on the structure and location of the pedicle relative to the two posterior openings. Kutorgina chengjiangensis, from the Chengjiang Lagerstätte of China, has a pedicle in the form of a thick annulated stalk. Despite its thickness, the pedicle is joined to the shell at the pseudodeltidium, strongly suggesting that it was originally derived from within the small pedicle foramen. The large posterior opening probably helped support strong diductor muscles in this interpretation. Each fossil includes a distinctive silicified tube within the large posterior opening. This tube has long been interpreted as a coprolite, suggesting that kutorginides had a complete gut terminating at a gap in the rear of the shell. This condition is otherwise unknown in articulate brachiopods, and is more akin to inarticulates in the subphylum Craniiformea. However, modern brachiopods have very small fecal pellets, so the silicified tube may be better interpreted as an adult pedicle. The smaller foramen at the tip of the pseudodeltidium may be a remnant of a larval pedicle, later rendered redundant by a second pedicle developing at the hinge in adulthood. In this scenario, the pedicle of other rhynchonelliforms would be homologous to the larval pedicle of Nisusia sulcata. ==Subgroups==
Subgroups
From The Treatise on Invertebrate Paleontology (Part H, Revised), unless stated otherwise: [mid-Cambrian (Wuliuan)] • Eoconcha Cooper, 1951 [lower-mid Cambrian] • Khasagtina? Ushatinskaya, 1987 [lower Cambrian ("Tommotian"? – "Atdabanian")] • Narynella Andreeva, 1987 [lower Cambrian ("Toyonian") – mid-Cambrian] • Nisusia Walcott, 1905 [lower Cambrian ("Botomian") – mid-Cambrian] • Trematosia Cooper, 1967 [lower Cambrian] • Incertae sedis genera: • Anomalocalyx Brock, 1999 [mid-Cambrian (Drumian)] == Representative genera ==
Representative genera
Kutorgina Some species of Kutorgina have a concavo-convex shell, with the smaller brachial (dorsal) valve dished in and the larger pedicle (ventral) valve broadly arched. The brachial valve has a rather prominent interarea at the back which is curved over by the prominent beak at the back of the pedicle valve. It includes the species Kutorgina elanica Malakhovskaya, 2013 and K. chengjiangensis Zhang et al. 2007, among many others. K. chengjiangensis preserves soft anatomy, including a pedicle, lophophore, and gut. Nisusia Nisusia Walcott, 1905 (Walcott, 1889) is known from the Miaolingian-age Burgess Shale (~). It is a senior synonym to Orthisina alberta Walcott, 1889. The pedicle of Nisusia emerges from between its valves, as displayed by silicified material of N. sulcata, though it still has an opening at the apex of the pedicle valve. ==References==
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