The genus
Lichinodium is
typified by
Lichinodium sirosiphoideum, first
described in 1875 by
William Nylander.
Aino Henssen added three species to the genus about a century later.
Lichinodium used to be classified in the
Lichinomycetes, but
molecular phylogenetic analysis published in 2018 showed that the genus was incorrectly classified there, and that instead it represents a new lichen-forming
lineage in the
superclass Sordariomyceta (containing the (
Leotiomycetes,
Laboulbeniomycetes, and the
Sordariomycetes), a major fungal group that was not previously known to have any lichenised species.
Lichinodium has a
sister taxon relationship with the family
Leotiaceae, and thus the order Lichinodiales is placed in the class
Leotiomycetes. Studies using
transmission electron microscopy also showed that in
Lichinodium, there are no
haustoria (microscopic root-like structures) made in the interface between mycobiont and cyanobiont; this difference further distinguishes
Lichinodium from the Lichinomycetes. ==Description==