, near
Ithaca, New York, in the
United States, an example of an upland river habitat In freshwater ecology, upland rivers and
streams are the fast-flowing rivers and streams that drain elevated or mountainous country, often onto broad alluvial plains (where they become lowland rivers). However, elevation is not the sole determinant of whether a river is upland or lowland. Arguably the most important determinants are those of
stream power and
stream gradient. Rivers with a course that drops rapidly in elevation will have faster water flow and higher stream power or "force of water". This in turn produces the other characteristics of an upland river—an
incised course, a
river bed dominated by
bedrock and coarse sediments, a
riffle and pool structure and cooler water temperatures. Rivers with a course that drops in elevation very slowly will have slower water flow and lower force. This in turn produces the other characteristics of a lowland river—a meandering course lacking
rapids, a river bed dominated by fine sediments and higher water temperatures. Lowland rivers tend to carry more suspended sediment and organic matter as well, but some lowland rivers have periods of high
water clarity in seasonal low-flow periods. The generally clear, cool, fast-flowing waters and bedrock and coarse sediment beds of upland rivers encourage fish species with limited temperature tolerances, high oxygen needs, strong swimming ability and specialised reproductive strategies to prevent eggs or larvae being swept away. These characteristics also encourage invertebrate species with limited temperature tolerances, high oxygen needs and ecologies revolving around coarse sediments and interstices or "gaps" between those coarse sediments. The term "upland" is also used in
wetland ecology, where "upland" plants indicate an area that is not a wetland. == Lowlands ==