The larvae are aquatic. The first and second
instar larvae probably breathe using both gills and
spiracles. They mainly crawl on substrates and sometimes back swim. The third to sixth instars do not have gills and use their spiracles to breathe. They back swim, meaning that they swim upside down. Larvae eat aquatic snails such as
Stenothyra formosana,
Gyraulus spirillus,
Gyraulus convexiusculus and
Lymnaea stagnalis. Predators of the larvae include the
red swamp crayfish,
Chinese mitten crab and
grass carp. In
Hubei, larvae overwinter at the bottom of the water. The pupae, about long, are light yellow, with some parts beginning to darken after three days. The pupae are enclosed in pupal cells made of earth. Adults live for about nine days, consuming only water. Mating takes several hours, after which the males quickly die. The eggs, orange-yellow in colour, are laid in masses on plants slightly below the water surface. Larvae emerge after about eleven days. ==References==