Mastocarpus papillatus

Despite the common name, it grows nowhere near Turkey. It is instead native to the northeastern Pacific Ocean and commonly can be found in California, Oregon, Washington, British Columbia, and Alaska. It has been found growing as far north as the Commander Islands and the Russian Far East, and as far south as Chile. ==Habitat==

It grows on rocky substrates in the high- to mid-intertidal zones and has been found growing at depths of 250 m. Like many red algae species, it uses phycoerythrin to photosynthesize which allows it to grow deeper than plants using other pigments. ==Description==

Turkish washcloth is reddish-brown to almost black, with males paler than females and lacking the signature papillae on the blades. Mazzaella affinis thalli looks similar in shape and color to male M. papillatus, but are smaller and grow in thick mats and not as individuals. It also can be confused with other species with Petrocelis phases , such as Pikea, other Mastocarpus spp., Ralfisa, and Hildenbrandia, thought the last two are thinner than the Turkish towel crust. ==Life cycle==

M. papillatus has a relatively complex reproductive cycle. Male gametophytes discharge nonflagellated sperm to drift in the current until they attach to the trichogynes of female plants. There the sperm perform mitosis without cell division, turning into a spermatium. Fertilisation then proceeds through a fertilization pore between the trichogyne and the spermatium. The eggs then grow into tetrasporophytes which are more or less copies of their parents. Alternately, the tetrasporophyte goes through the Petrocelis phase (so named as it was thought to be a distinct genus, Petrocelis, of Phyllophoraceae) and grows into a crust that looks like spilled tar, inspiring the black tar spot common name. The tetrasporophyte undergoes meiosis and produce tetrasporangia which release spores to germinate elsewhere. Eventually the crust grows into a typical adult plant. This crustose phase is shared by all members of the genus Mastocarpus as well as those in Pikea. ==Ecology==

Turkish washcloth is food for many animals, including Echinolittorina ziczac, Littorina, Amphipoda, Lottia, Pholis crassispina, Lottia scutum, Trochidae, Lacuna vincta, Stenosoma, Mopalia swanii, and Cryptochiton stelleri. Like the Porphyra species used to make nori, M. papillatus is susceptible to infection by the parasitic oomycete Pythium porphyrae, or red rot disease, which can kill large colonies. ==Uses==

Grapestone is edible Grapestone is easily foraged from the shore, but like any wild species overharvesting is possible. By grabbing the longer blades and pulling sideways, the shorter blades and holdfast may remain on the rock and live on. M. papillatus is also thought to have antimicrobial properties. ==Gallery==

File:Mastocarpus papillatus, Cambria.jpg File:Mastocarpus papillatus from Vancouver.jpg File:Mastocarpus papillatus on Petrocelis.jpg == References ==